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Main page: Category:Campanian life

During the Campanian age, a radiation among dinosaur species occurred. In North America, for example, the number of known dinosaur genera rises from 4 at the base of the Campanian to 48 in the upper part. This development is sometimes referred to as the "Campanian Explosion". However, it is not yet clear if the event is artificial, i.e. the low number of genera in the lower Campanian can be caused by a lower preservation chance for fossils in deposits of that age. The generally warm climates and large continental area covered in shallow sea during the Campanian probably favoured the dinosaurs. In the following Maastrichtian stage, the number of North American dinosaur genera found is 30% less than in the upper Campanian.

Animals that lived in the Campanian include:

Cartilaginous fish

Cartilaginous fish of the Campanian
Taxa	Presence	Location	Description	Images
Chlamydoselachus				Chlamydoselachus
†Schizorhiza

Actinopterygii

Ray-finned fish of the Campanian
Taxa	Presence	Location	Description	Images
†Gillicus	100–66 Ma	North America	A genus of osteoglossomorph fish in the family Ichthyodectidae. Unlike its larger relative, Xiphactinus, by which it was preyed upon, it was a filter feeder instead of a large prey predator.	Gillicus Ichthyodectes Xiphactinus
†Ichthyodectes	Coniacian to Campanian	North America	A genus of large osteoglossomorph fish in the family Ichthyodectidae. Like its larger close relative Xiphactinus was a voracious predator.
†Xiphactinus	Albian to Maastrichtian	North America; Europe; Australia; Venezuela	A genus of large osteoglossomorph fish in the family Ichthyodectidae that could reach up to 4 meters in length. Xiphactinus was a voracious predator.

Actinistia

Coelacanths of the Campanian
Taxa	Presence	Location	Description	Images
†Axelrodichthys †Axelrodichthys megadromos	Lower Campanian possibly to early Maastrichtian	France	A species of averagely sized mawsoniid coelacanth. Related species of the same genus lived mostly on earlier stages of the Cretaceous.	Megalocoelacanthus
†Megalocoelacanthus Megalocoelocanthus dobiei	Lower Campanian possibly to early Maastrichtian	USA	A very large latimerid coelacanth. It has been estimated to have been 3.5 - 4.5 meters in length.

Dipnoi

Lungfish of the Campanian
Taxa	Presence	Location	Description	Images
†Ceratodus	Olenekian-Eocene	Cosmpopolitan	A widespread genus of ceratodontiform lungfish in the family Ceratodontidae with multiple species throughout the mesozoic.	Ceratodus Retodus
Protopterus	Campanian to Recent	Egypt, Sudan	A ceratodontiform lungfish in the family Protopteridae. Species of this genus, survive to this day.
†Retodus	Albian–Maastrichtian	Egypt, Sudan	A genus of large ceratodontiform lungfish in the family Ceratodontidae.

Lepidosaurs

Lepidosaurs of the Campanian
Taxa	Presence	Location	Description	Images
†Adamisaurus †A. magnidentatus			An agamid iguanian lizard.	Clidastes Ectenosaurus Estesia Globidens alabamaensis Halisaurus arambourgi Mosasaurus beaugei Mosasaurus lemonnieri Mosasaurus missouriensis Platecarpus tympaniticus Prognathodon lutugini Taniwhasaurus Telmasaurus Tylosaurus pembinensis Tylosaurus proriger
†Aiolosaurus †A. oriens			A varanoid lizard of disputed affinity.
†Alamitophis †A. argentinus †A. elongatus			A genus of fossil snakes in the extinct family of Madtsoiidae.
†Anchaurosaurus		Djadochta Formation, Gobi Desert, Mongolia	An iguanian lizard in the extinct clade Gobiguania. Compared to other iguanians, Anchaurosaurus has a relatively elongated skull, large eye sockets, and higher tooth crowns.
†Apsgnathus		Brewster County, Aguja Formation, Texas, USA	A scincomorph.
†Asprosaurus			A genus of anguimorph lizard distantly related to the living Gila monster.
†Bainguis	84.9-70.6 mya	Bayan Mandahu Formation, Mongolia	A diploglossan lizzard.
†Catactegenys		Brewster County, Aguja Formation, Texas, USA	A large, possibly durophagous, night lizard.
†Carusia			A basal anguimorph lizard closely related to the extant Xenosauridae.
†Cherminotus			A varanoid lizard that resembles Aiolosaurus.
†Clidastes			A relatively small basal mosasaurine marine lizard.
†Ctenomastax		Djadochta Formation, Gobi Desert, Mongolia	An iguanid iguanian lizard.
†Eoxanta		Djadochta Formation, Gobi Desert, Mongolia	A scincomorph lizard.
†Dryadissector †D. shilleri			A genus of extinct varanoid lizards.
†Ectenosaurus			A genus of plioplatecarpine mosasaurs.
†Eonatator †E. coellensis †E. sternbergii			A genus of halisaurine mosasaurs.
†Estesia		Gobi Desert, Mongolia	A genus of anguimorph lizards, originally thought to be closely related to monitor lizards.
†Flaviagama		Djadochta Formation, Gobi Desert, Mongolia	A priscagamid iguanian lizard.
†Globaura		Djadochta Formation, Gobi Desert, Mongolia	A scincomorph lizard.
†Gobiderma		Djadochta Formation, Gobi Desert, Mongolia	An anguimorph lizard that resembled a Gila monster though it was not necessarily closely related to it.
†Gobinatus		Djadochta Formation, Gobi Desert, Mongolia	A teiid lacertoid lizard.
†Globidens †G. alabamaensis †G. dakotensis †G. hisaensis †G. schurmanni †G. simplex			A genus of medium sized mosasaurine mosasaurs. Along with its closest relatives in the tribe Globidensini it had distinctive globular teeth.
†Gobiderma			An anguimorph lizard that resembled the modern Heloderma lizards.
†Halisaurus			A comparatively small mosasaurs. Type genus of the subfamily Halisaurine.
†Herensugea			A madtsoiid snake.
†Hymenosaurus		Djadochta Formation, Gobi Desert, Mongolia	A scincomorph lizard.
†Isodontosaurus		Djadochta Formation, Gobi Desert, Mongolia	An iguanian lizard in the extinct clade Gobiguania.
†Kawasphenodon †K. expectatus			A sphenodontid rhynchocephalian closely related to the extant tuatara.
†Lamarquesaurus †L. cabazai			A basal neosphenodont rhynchocephalian.
†Latoplatecarpus †L. nichollsae †L. willistoni		New Zealand, Japan, Antarctica	A large plioplatecarpine mosasaur.
†Menarana †M. laurasiae			A madtsoiid snake.
†Mimeosaurus			A priscagamid iguanian lizard. Mimeosaurus is unique among iguanians in having premaxilla bones at the tip of the snout that are reduced in size, as well as having two pairs of enlarged canine-like teeth in the maxilla
†Mosasaurus †M. beaugei †M. conodon †M. lemonnieri †M. missouriensis			A genus of large mosasaurine mosasaurs, contained some of the largest marine reptiles to have ever lived.
†Myrmecodaptria		Djadochta Formation, Gobi Desert, Mongolia	A basal lizard of contested affinities. Might be related to the contemporary Bainguis.
†Odaxosaurus †O. piger †O. priscus			An anguid anguimorph.
†Ovoo †O. gurvel'			An early varanid.
†Paravaranus †P. angustifrons			A varanoid lizard.
†Parmeosaurus		Djadochta Formation, Gobi Desert, Mongolia	A scincomorph lizard.
†Patagoniophis †P. parvus			A madtsoiid snake.
†Phrynosomimus		Djadochta Formation, Gobi Desert, Mongolia	A priscagamid iguanian lizard.
†Platecarpus		USA, possibly Belgium and Africa	A genus of plioplatecarpine mosasaurs.
†Pleurodontagama		Bayan Mandahu Formation Gobi Desert, Mongolia	Originally thought to be a priscagamid, it's sub-pleurodont dentition instead of acrodont permanent dentition indicates that it might had been a more basal iguanian, a potential "missing link" between Chamaeleontiformes and Pleurodonta.
†Plioplatecarpus †P. houzeaui †P. peckensis †P. primaevus			A genus of plioplatecarpine mosasaurs.
†Priscagama		Djadochta Formation, Gobi Desert, Mongolia	A priscagamid iguanian lizard.
†Prognathodon †P. currii †P. lutugini †P. overtoni			A genus of large mosasaurine mosasaurs.
†Proplatynotia †P. proplatynotia			An anguimorph lizard closely related to Monstersauria Varanoidea.
†Rionegrophis			A madtsoiid snake.
†Saichangurvel		Djadochta Formation, Gobi Desert, Mongolia	An iguanian lizard in the extinct clade Gobiguania.
†Saniwides			A varanoid lizard.
†Slavoia		Djadochta Formation, Gobi Desert, Mongolia	Previously thought to be a scincomorph, it was later placed in a clade containing Lacertoidea+Scincomorpha. A more recent description of the genus, placed it as a stem-group relative of amphisbaenians within Lacertoidea.
†Taniwhasaurus		New Zealand, Japan, Antarctica	A medium sized tylosaurine mosasaur.
†Temujinia		Djadochta Formation, Gobi Desert, Mongolia	An iguanian lizard in the extinct clade Gobiguania.
†Telmasaurus			A varanoid lizard.
†Tylosaurus †T. bernardi †T. ivoensis †T. pembinensis †T. proriger †T. saskatchewanensis			A genus of large tylosaurine mosasaurs, contained some of the largest marine reptiles to have ever lived.
†Zapsosaurus		Djadochta Formation, Gobi Desert, Mongolia	An iguanian lizard in the extinct clade Gobiguania.

Testudines

Turtles of the Campanian
Taxa	Presence	Location	Description	Images
†Allopleuron				Archelon Protostega
†Archelon
†Atlantochelys
†Calcarichelys
†Chelosphargis
†Ctenochelys
†Desmatochelys
†Leviathanochelys
†Mesodermochelys
†Nichollsemys
†Pneumatoarthrus
†Protostega
†Shandongemys			A lindholmemydid pantestudinoid turtle.
†Terlinguachelys
†Toxochelys

†Plesiosaurs

Plesiosaurs of the Campanian
Taxa	Presence	Location	Description	Images
†Elasmosaurus	80.5 mya	Pierre Shale, Kansas, USA	Elasmosaurus is a genus of plesiosaur with an extremely long neck.	Elasmosaurus Styxosaurus
†Styxosaurus	83.5-80.5 mya	Logan County, Kansas	Styxosaurus is an Elasmosaurid plesiosaur.

†Choristoderans

Choristoderans of the Campanian
Taxa	Presence	Location	Description	Images
†Champsosaurus	Campanian to Paleocene	North America	Champsosaurus is a genus of crocodile-like neochoristodere.	Champsosaurus natator

Crocodylomorphs

Crocodylomorphs of the Campanian
Taxa	Presence	Location	Description	Images
†Acynodon †Acynodon adriaticus †Acynodon iberoccitanus	Early Campanian - Late Maastrichtian 83.5–66 Ma	Italy	Initially placed within Alligatoridae but has since been reclassified as a more basal globidontan, the oldest and most primitive known to date.	Allodaposuchus precedens Brachychampsa montana Deinosuchus riograndensis Deinosuchus rugosus Paralligator gradilifrons
†Albertochampsa †Albertochampsa langstoni	76.9–75.8 Ma	Dinosaur Park Formation, Alberta	A basal member of Globidonta, a group of alligatoroids more derived than Deinosuchus or Leidyosuchus that includes living alligators and caimans.
†Agaresuchus †Agaresuchus fontisensis	Late Campanian-Maastrichtian	Spain	A genus of average-sized basal eusuchian. Might be synonymous with its contemporary relative genus Allodaposuchus.
†Allodaposuchus †Allodaposuchus precedens †Allodaposuchus iberoarmoricanus	84.9–66.043 Ma	Spain and France,	A genus of average-sized basal eusuchian, growing to around 3 m (9.8 ft) long. The main feature that distinguishes this species from other related crocodylomorphs is the orientation of a groove at the back of the skull called the cranioquadrate passage; unlike the cranioquadrate passages of other crocodylomorphs, which are only visible at the back of the skull, the cranioquadrate passage of this variety is visible when the skull is viewed from the side. The genus includes many species but it may be paraphyletic in regards to its contemporary relative genera Agaresuchus and Lohuecosuchus.
†Artzosuchus †Artzosuchus brachicephalus	Campanian	Djadochta Formation, Mongolia	Its classification beyond that of a basal crocodylomorph is indeterminant because of the fragmentary nature of the material associated with the genus.
†Brachychampsa †Brachychampsa montana †?Brachychampsa sealeyi	early Campanian to Danian	Hell Creek Formation, Montana, USA	A globidontan distinguished by an enlarged fourth maxillary tooth in the upper jaw.
†Deinosuchus †Deinosuchus hatcheri †Deinosuchus riograndensis †Deinosuchus rugosus †Deinosuchus schwimmeri	82–73 Ma	USA	A basal alligatoroid and one of the largest crocodylomorphs that ever lived.
†Gobiosuchus †Gobiosuchus kielanae	Early Campanian 75–71 Ma	Djadokhta Formation, Gobi Desert, Mongolia.	A member of Gobiosuchidae, a family of late surviving but basal non-mesoeucrocodilian crocodyliforms.
†Leidyosuchus †Leidyosuchus canadensis	Middle Campanian	Alberta	A basal medium-sized alligatoroid.
†Lohuecosuchus †Lohuecosuchus megadontos	Late Campanian-early Maastrichtian	Spain	A genus of average-sized basal eusuchian. Might be synonymous with its contemporary relative genus Allodaposuchus.
†Paralligator †Paralligator gradilifrons	Late Cretaceous, ~96–70 Ma	Nemegt Formation, Mongolia	A paralligatorid mesoeucrocodylian.
†Shamosuchus †Shamosuchus djadochtaensis	83.6–72.1 mya	Djadochta Formation, Mongolia	A paralligatorid mesoeucrocodylian.
†Zaraasuchus †Zaraasuchus shepardi		Zos Canyon, Gobi Desert, Mongolia.	A gobiosuchid.

†Pterosaurs

Pterosaurs of the Campanian
Taxa	Presence	Location	Description	Images
Aerotitan	Campanian-Maastrichtian	Allen Formation, Patagonia, Argentina	A genus of moderately large azhdarchid pterosaurs.	Aerotitan Nyctosaurus Pteranodon
Bogolubovia		Rybushka Formation, Petrovsk, Russia	A genus of moderately of medium to large sized azhdarchid pterosaurs.
Geosternbergia		USA, North America	Geosternbergia was originally a species of Pteranodon and is famous for its oddly shaped crest.
Montanazhdarcho		Montana, USA	Small azhdarchoid pterosaur, probably a tapejarid
Navajodactylus		New Mexico, USA, and Alberta, Canada	Known primarily from forearm elements; tentatively assigned to Azhdarchidae, though most likely not part of it.
Nyctosaurus		mid-western United States	Nyctosaurus is a genus of pterodactyloid pterosaur.
Piksi		Montana, USA	Piksi is a genus of pterosaurs containing the single species Piksi barbarulna.
Pteranodon		Kansas, USA, North America	Pteranodon is a genus of pterosaurs which included some of the largest known flying reptiles, with wingspans over 6 metres
Volgadraco		Saratov, Russia	A large pterosaur, either an azhdarchoid or a nyctosaurid.

†Dinosaurs

†Ornithischia

†Ankylosaurs

Ankylosaurs of the Campanian
Taxa	Presence	Location	Description	Images
Aletopelta		Point Loma Formation, California, USA	A medium-sized ankylosaurid, estimated to be around 6 m (20 ft) long.	Edmontonia Euoplocephalus Pinacosaurus
Antarctopelta		Santa Marta Formation, James Ross Island, Antarctica	A stocky ankylosaur protected by armor plates embedded in the skin. Although a complete skeleton has not been found, the species is estimated to have reached a maximum length of 4 meters (13 feet). Displays characteristics of both ankylosaurids and nodosaurids.
Edmontonia	Campanian to Maastrichtian	Horseshoe Canyon Formation, Alberta, Canada	A bulky nodosaurid at roughly 6.6 m (22 ft) long. It had small, ridged bony plates on its back and many sharp spikes along its body sides. The four largest spikes jutted out from the shoulders on each side, two of which were split into subspines in some specimens. Its skull had a pear-like shape when viewed from above.
Euoplocephalus
Heishansaurus			A dubious ankylosaurid, possibly a synonym of Pinacosaurus.
Nodocephalosaurus
Palaeoscincus		Judith River Formation	known from a single tooth
Panoplosaurus		Judith River Formation, Alberta, Canada; Montana, USA	A 5.5–7 m long nodosaurid.
Pinacosaurus
Saichania
Shanxia
Struthiosaurus
Tarchia
Tianzhenosaurus

†Theskelosaurids

Parksosaurids of the Campanian
Taxa	Presence	Location	Description	Images
Orodromeus	76.7 mya	Montana, USA	Orodromeus was a small fast bipedal herbivore that probably coexisted with dinosaurs such as Daspletosaurus and Einiosaurus. Its length was estimated by Horner & Weishampel at 2.5 metres.	Orodromeus

†Ceratopsians

Ceratopsians of the Campanian
Taxa	Presence	Location	Description	Images
Achelousaurus Achelousaurus horneri	74.2 million years ago	Glacier County, Two Medicine Formation, Montana, U.S.A.	A centrosaurine.	Achelousaurus Agujaceratops Albertaceratops Anchiceratops Avaceratops Bagaceratops Brachyceratops Breviceratops Centrosaurus Cerasinops Chasmosaurus Coahuilaceratops Coronosaurus Diabloceratops Einiosaurus Graciliceratops Judiceratops Kosmoceratops Lamaceratops Medusaceratops Micropachycephalosaurus Mercuriceratops Nasutoceratops Pachyrhinosaurus lakustai Pentaceratops Prenoceratops Protoceratops Rubeosaurus Spinops Styracosaurus Titanoceratops Udanoceratops Utahceratops Vagaceratops Xenoceratops Yehuecauhceratops
Agujaceratops Agujaceratops mariscalensis Agujaceratops mavericus	77 million years ago	Aguja Formation, Texas	A chasmosaurine.
Albertaceratops Albertaceratops nesmoi
Anchiceratops Anchiceratops longirostris
Avaceratops Avaceratops lammersi
Bagaceratops Bagaceratops rozhdestvenskyi
Bainoceratops Bainoceratops efremovi
Brachyceratops Brachyceratops montanensis
Breviceratops Breviceratops kozlowskii
Centrosaurus Centrosaurus apertus
Cerasinops Cerasinops hodgskissi
Ceratops Ceratops montanus
Chasmosaurus Chasmosaurus russelli Chasmosaurus belli
Coahuilaceratops Coahuilaceratops magnacuerna
Coronosaurus Coronosaurus brinkmani
Diabloceratops Diabloceratops eatoni
?Dysganus Dysganus encaustus Dysganus bicarinatus Dysganus peiganus
Einiosaurus Einiosaurus procurvicornis
Eoceratops Eoceratops canadensis
Graciliceratops Graciliceratops mongoliensis
Gryphoceratops Gryphoceratops morrisoni
Judiceratops Judiceratops tigris
Kosmoceratops Kosmoceratops richardsoni
Lamaceratops Lamaceratops tereschenkoi
Magnirostris Magnirostris dodsoni
Medusaceratops Medusaceratops lokii
Mercuriceratops Mercuriceratops gemini
Micropachycephalosaurus Micropachycephalosaurus hongtuyanensis
Mojoceratops Mojoceratops perifania
Monoclonius Monoclonius crassus
Nasutoceratops Nasutoceratops titusi
?Notoceratops Notoceratops bonarellii		Chubut Province, Argentina	A dubious genus of possible ceratopsian affinity
Pachyrhinosaurus Pachyrhinosaurus lakustai
Pentaceratops Pentaceratops sternbergii
Platyceratops Platyceratops tatarinovi
Prenoceratops Prenoceratops pieganensis
Protoceratops Protoceratops andrewsi Protoceratops hellenikorhinus
Rubeosaurus Rubeosaurus ovatus
Spiclypeus Spiclypeus shipporum
Spinops Spinops sternbergorum
Styracosaurus Styracosaurus albertensis
Titanoceratops Titanoceratops ouranos
Udanoceratops Udanoceratops tschizhovi
Unescoceratops Unescoceratops koppelhusae
Utahceratops Utahceratops gettyi
Vagaceratops Vagaceratops irvinensis
Xenoceratops Xenoceratops foremostensis
Yehuecauhceratops Yehuecauhceratops mudei	72 mya	Coahuila, Aguja Formation, Mexico	A centrosaurine.

†Pachycephalosaurs

Pachycephalosaurs of the Campanian
Taxa	Presence	Location	Description	Images
Colepiocephale		Alberta, Canada	The oldest known pachycephalosaurid.	Colepiocephale lambei Goyocephale Hanssuesia sternbergi Homalocephale Prenocephale Sphaerotholus Stegoceras Tylocephale Wannanosaurus yansiensis
Goyocephale		Mongolia
Gravitholus
Hanssuesia		Alberta, Canada; Montana, USA	Distinguished from other pachycephalosaurs by having a depressed parietal region, wide frontoparietal dome, broad nasal characteristics, reduced prefontal lobes, and a reduced parietosquamosal shelf.
Homalocephale		Mongolia	Sporting a flat, wedge-shaped skull roof, Homalocephale was different from other pachycephalosaurs.
Ornatotholus			A possible synonym of [Stegoceras]].
Prenocephale
Sphaerotholus
Stegoceras
Tylocephale
Texacephale	77 mya	Aguja Formation, Texas
Wannanosaurus

†Ornithopods

Ornithopods of the Campanian
Taxa	Presence	Location	Description	Images
Acristavus	79 mya	Montana and Utah, USA	A genus of basal saurolophine hadrosaur.	Acristavus gagslarsoni Angulomastacator daviesi Aralosaurus tuberiferus Brachylophosaurus Corythosaurus casuarius Edmontosaurus regalis Gasparinisaura Gryposaurus notabilis Hadrosaurus foulkii Hypacrosaurus altispinus Kritosaurus Lambeosaurus lambei Malefica Maiasaura Naashoibitosaurus ostromi Nipponosaurus Ornatops Parasaurolophus walkeri Prosaurolophus Rhabdodon Saurolophus angustirostris Shantungosaurus Tanius Tsintaosaurus Velafrons
Angulomastacator	76.9 mya	Aguja Formation, Texas	A lambeosaurine.
Aquilarhinus		Aguja Formation, Texas	Formerly referred to Kritosaurus and Gryposaurus it might be a much more basal hadrosaurid.
Aralosaurus	Upper Santonian-Lower Campanian boundary 85.8 mya	Bostobe Formation, Kazakhstan	A basal lambeosaurine hadrosaur, Aralosaurus was about the size of an elephant. Although very little is known about Aralosaurus (only one near complete skull has been found); it was identified by a beak with nearly 1,000 small teeth in 30 rows. The back of an Aralosaurus skull was wide, a feature suggestive of large jaw muscles used to power its chewing apparatus.
Brachylophosaurus	76.5 mya	Montana, USA; Alberta, Canada	Brachylophosaurus was a basal saurolophine hadrosaur which reached an adult length of 9 meters (30 feet).
Corythosaurus	77-76.5 mya	Alberta, Canada	A derived lambeosaurine. Corythosaurus weighed in at 4 tonnes and measured roughly 10 metres (33 feet) from nose to tail.
Diclonius	75 mya	Montana, USA	A dubious genus of hadrosaur known only from dental fossils.
Edmontosaurus	73.0-76.5 mya	Canada	Edmontosaurus, a saurolophine hadrosaur, included some of the largest ornithischian species, measuring up to 12 metres (39 feet) long and weighing around 4.0 metric tons (4.4 short tons).
Gasparinisaura	85 mya	Argentina	Gasparinisaura was a small ornithopod of the basal clade Elasmaria. In 2010 Gregory S. Paul estimated the length at 1.7 metres, the weight at thirteen kilogrammes.
Gilmoreosaurus	72 mya	Mongolia	A relatively large hadrosauromorph.
Gryposaurus	83-75.5 mya	Alberta, Canada	Gryposaurus was a saurolophine hadrosaur of typical size and shape.
Hadrosaurus	79.5 mya	New Jersey, USA	One of the most basal hadrosaurids and the namesake genus of the family. It was likely bipedal for the purposes of running, but could use its forelegs to support itself while feeding.
Hypacrosaurus	75-67 mya	Alberta, Canada	A derived lambeosaurine, Hypacrosaurus is most easily distinguished from other hollow-crested duckbills by its tall neural spines and the form of its crest. The neural spines, which project from the top of the vertebrae, are 5 to 7 times the height of the body of their respective vertebrae in the back,[4] which would have given it a tall back in profile. The skull's hollow crest is like that of Corythosaurus, but is more pointed along its top, not as tall, wider side to side, and has a small bony point at the rear
Hypsibema		North Carolina and Missouri, USA	A large hadrosauroid known only by a couple of vertebrae.
Kritosaurus	73 mya	North America	The type specimen of Kritosaurus navajovius is only represented by a partial skull and lower jaws, and associated postcranial remains.
Lambeosaurus	76-75 mya	Alberta, Canada	A derived lambeosaurine hadrosaur, namesake genus of the subfamily.
Lophorhothon	80 mya	Alabama, USA	A genus of hadrosauromorphs.
Malefica		Aguja Formation, Texas	A basal hadrosaur.
Maiasaura	76.7 mya	Montana, USA	Maiasaura was a large basal saurolophine hadrosaur, attaining an adult length of about 9 metres (30 feet) and had the typical hadrosaurid flat beak and a thick nose. It had a small, spiky crest in front of its eyes. The crest may have been used in headbutting contests between males during the breeding season.
Microhadrosaurus		China	A possibly dubious genus of hadrosaur known by a partial lower jaw from a juvenile.
Mochlodon		Austria	A rhabdodontid.
Naashoibitosaurus	73 mya	New Mexico, USA	Naashoibitosaurus, a saurolophine hadrosaur, based as it is on a single partial skeleton, is not well known in terms of anatomy. Its skull, the most thoroughly described portion, has a low nasal crest that peaks in front of the eyes, but does not strongly arch as in Gryposaurus.
Nipponosaurus	80 mya	Sakhalin island	A lambeosaurine hadrosaur known from subadult remains.
Ornatops	78 mya	New Mexico, USA	A genus of basal saurolophine hadrosaur.
Parasaurolophus	76.5-73 mya	Alberta, Canada; New Mexico and Utah, USA	A large derived lambeosaurine hadrosaur.
Probrachylophosaurus	79 mya	Montana, USA; Alberta, Canada	A large basal saurolophine hadrosaur which reached an adult length of over 9 meters (30 feet).
Prosaurolophus	76-75 mya	Alberta, Canada	Prosaurolophus was a large-headed duckbill; the most complete described specimen has a skull around 0.9 meters (3.0 feet) long on a ~8.5 meter long skeleton (~28 ft).[2] It had a small, stout, triangular crest in front of the eyes; the sides of this crest were concave, forming depressions. The upper arm was relatively short.
Pteropelyx	75 mya	Montana, USA	A dubious genus of lambeosaurine hadrosaurs. Historically, several species were assigned to it, all based on extremely fragmentary remains, but there is no evidence to support these assignments.
Rhabdodon	72 mya	France; Spain; Haţeg Island, Romania	It is unclear whether it was an iguanodont or a hypsilophodont, and may be a "missing link" between the two. Current evidence indicates it is an iguanodont similar to Tenontosaurus.
Saurolophus	69.5-68.5 mya	North America, Asia	Saurolophus is known from material including nearly complete skeletons, giving researchers a clear picture of its bony anatomy. S. osborni, the rarer Albertan species, was around 9.8 meters (32 feet) long, with its skull a meter long (3.3 feet). Its weight is estimated at 1.9 tonnes (2.1 tons). S. angustirostris, the Mongolian species, was larger; the type skeleton is roughly 12 meters (39 feet) long, and larger remains are reported.
Shantungosaurus	72 mya	China	A very large saurolophine hadrosaur closely related to Edmontosaurus, it is one of the longest and largest known hadrosaurids and possibly the largest non-sauropod dinosaur; the composite skeleton of a medium-sized individual mounted at the Geological Institute of China in Beijing measures 14.72 metres (48.3 feet) in length.
Stephanosaurus	76–75 mya	Dinosaur Park Formation, Alberta	A dubious genus of hadrosaurid dinosaur with a complicated taxonomic history.
Tanius	72–71 mya	China	A genus of hadrosauromorph dinosaur.
Trachodon	77 mya	Montana, USA	Trachodon is a dubious genus of hadrosaurid dinosaur based on teeth. Numerous species had been referred to this genus later reassigned to different genera or proven dubious.
Tsintaosaurus	72 mya	Southern China	A basal but relatively large lambeosaurine hadrosaur.
Velafrons	72 mya	Mexico	A large derived lambeosaurine.

†Sauropoda

Sauropods of the Campanian
Taxa	Presence	Location	Description	Images
Alamosaurus		Southwestern United States	Alamosaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now North America. It was a large quadrupedal herbivore.	Alamosaurus Andesaurus Dreadnoughtus Huabeisaurus allocotus Neuquensaurus Overosaurus Quaesitosaurus Saltasaurus
Andesaurus		Neuquén Province, Argentina	Andesaurus is a genus of basal titanosaurian sauropod dinosaur.
Argyrosaurus		Lago Colhué Huapi Formation, Argentina
Dreadnoughtus		Cerro Fortaleza Formation, Argentina	Dreadnoughtus is one of the largest titanosaurs known.
Gondwanatitan		Adamantina Formation and Cambabe Formation, Brazil
Huabeisaurus		North East, China	A member of the Euhelopodidae sauropods.
Laplatasaurus		Allen Formation and Anacleto Formation, both in Argentina; Palacio Formation, Uruguay
Loricosaurus	Campanian-Maastrichtian	Allen Formation, Argentina
Microcoelus	Santonian-Campanian	Bajo de la Carpa Formation, Argentina
Neuquensaurus		Anacleto Formation, Argentina
Overosaurus		Neuquén Province, Argentina	Small-sized titanosaur.
Quaesitosaurus		Shar Tsav, Mongolia	Quaesitosaurus is a genus of titanosaurian sauropod.
Saltasaurus		north-west Argentina; Uruguay	Saltasaurus is a genus of titanosaurid sauropod dinosaur. An estimated length of 12 metres (39 feet) and a mass of 7 tonnes (8 tons).
Rocasaurus	Campanian-Maastrichtian	Allen Formation, Rio Negro Province, Argentina

Theropoda

David J. Varrichio observes that during the late Campanian Alberta and Montana had very similar theropods despite significant differences in the types of herbivorous dinosaur faunas.

Non-avian theropods of the Campanian
Taxa	Presence	Location	Description	Images
†Abelisaurus		Allen Formation?, Anacleto Formation?, Argentina	An abelisaurid ceratosaur that probably reached 7 to 9 meters in length; known from only one partial skull.	Abelisaurus Appalachiosaurus Archaeornithomimus Bambiraptor Carnotaurus Citipati Daspletosaurus Dromaeosaurus Gobivenator Mahakala Oviraptor Pyroraptor Saurornithoides Shuvuuia Struthiomimus altus Teratophoneus Tsaagan Velociraptor
†Appalachiosaurus			A derived tyrannosauroid.
†Archaeornithomimus			An ornithomimid ornithomimosaur.
†Bambiraptor			A small dromaeosaurid.
†Bistahieversor			A large derived tyrannosauroid.
†Byronosaurus			A troodontid
†Citipati			An oviraptorid oviraptorosaur.
†Carnotaurus			A derived abelisaurid ceratosaur that probably reached 7 to 9 meters in length
†Chirostenotes			A caenagnathid oviraptorosaur.
†Daspletosaurus †Daspletosaurus torosus †Daspletosaurus wilsoni †Daspletosaurus horneri †Daspletosaurus sp.			A genus of large and robustly built tyrannosaurines.
†Deinodon		Judith River Formation	A dubious tyrannosaurid known from tooth remains.
†Dromaeosaurus			A small to medium sized dromaeosaurid maniraptoran theropod with a sickle-like claw.
†Dromiceiomimus			An ornithomimid ornithomimosaur.
†Dryptosaurus			A derived tyrannosaur.
†Dynamoterror			A large basal tyrannosaurine.
†Gobivenator			A troodontid
†Gorgosaurus			An albertosaurine tyrannosaurid.
†Harpymimus			A basal ornithomimosaur.
†Khaan			An oviraptorid oviraptorosaur.
†Kol			An alvarezsaurid.
†Labocania			A large robust theropod of uncertain affinities, possibly a tyrannosaur.
†Leptorhynchos			A caenagnathid oviraptorosaur.
†Linheraptor			A dromaeosaurid.
†Lythronax		Wahweap Formation, Utah	A 7-meter tyrannosaurine known from a partially complete skull, some vertebrae and a complete pubis
†Luanchuanraptor			A dromaeosaurid.
†Mahakala			A small unenlagiid maniraptoran.
†Nanshiungosaurus			A relatively large-bodied therizinosaurid.
†Noasaurus			A small derived ceratosaur.
†Ornithomimus			An ornithomimid ornithomimosaur.
†Oviraptor		Mongolia	Oviraptor is an oviraptorid oviraptorosaur in the subfamily Oviraptorinae.
†Parvicursor			An alvarezsaur.
†Pyroraptor		Var, France	A small dromaeosaurid.
†Richardoestesia			A coelurosaur form taxon known only from teeth and lower jaw fragments.
†Saurornithoides			A troodontid.
†Saurornitholestes			A dromaeosaurid.
†Shuvuuia			An alvarezsaur.
†Struthiomimus			An ornithomimid ornithomimosaur.
†Teratophoneus			A basal tyrannosaurine.
†Thanatotheristes			A close relative of Daspletosaurus.
†Troodon			A troodontid.
†Tsaagan			A dromaeosaurid.
†Variraptor		Var, France	A dromaeosaurid.
†Velociraptor		Mongolia and China	A dromaeosaurid.
†Zhuchengtyrannus		Wangshi Group, Zhucheng, China	One of the largest tyrannosaurids at between 10–12 meters. Known from a lower jaw and maxilla slightly smaller than those of the later Tyrannosaurus.

Avialae

Birds of the Campanian
Taxa	Presence	Location	Description	Images
†Alexornis			A genus of enantiornithine birds of uncertain affinities.	Gargantuavis Hesperornis Hollanda Ichthyornis
†Apatornis			A genus of basal ornithuran birds.
†Apsaravis			A genus of basal ornithuromorph birds.
†Baptornis			A genus of flightless hesperornithiform birds.
†Brodavis			A genus of freshwater hesperornithiform birds. Although Hesperornithes probably lost their flight abilities by the end of the Early Cretaceous, minimal pachyostosis in Brodavis suggests the possibility of some volant abilities.
†Canadaga			A genus of flightless hesperornithid birds.
†Castignovolucris			An enantiornithe of uncertain affinities. It was estimated to have a wingspan of around 127–185 cm and may have been 75 to 110 cm long when fully grown, making it one of the largest known enantiornitheans to date.
†Cretaaviculus	~85 mya	Bostobe Formation, Kazakhstan	An extinct genus of birds. The type species, C. sarysuensis, is known only from an isolated, asymmetrical contour feather
†Elsornis			Elsornis is a genus of basal enantiornithine birds. It was flightless or nearly so.
†Gargantuavis			A genus of large, primitive and potentially flightless avialan.
†Gettyia			An avisaurid enantiornithe, previously thought to be a species of the later closely related Maastrichtian genus Avisaurus.
†Gobipteryx			A genus of euenantiornithine enantiornithe. It possessed a robust beak which convergently evolved with those of modern birds.
†Halimornis			A genus of avisaurid enantiornithe. It lived in a coastal environment.
†Hesperornis			A genus of flightless hesperornithid birds.
†Hollanda			A genus of small ground birds. It was either an enantiornithe or a basal ornithuromorph.
†Ichthyornis			A genus of toothy seabird-like ornithuran.
†Intiornis			A genus of avisaurid enantiornithe. Although closely related to some of the largest avisaurids, members of this genus were very small birds.
†Maaqwi			A genus of large marine diving vegaviid bird with an estimated body mass of 1.5 kg.
†Mirarce			A genus of avisaurid enantiornithes, closely related to the later genus Avisaurus.
†Limenavis			Limenavis is the closest known relative of modern birds.
†Neogaeornis wetzeli			A marine bird from Chile. It had the midfeet of a foot-propelled diving bird, but its relationships are enigmatic. The only known species is from the Campanian-Maastrichtian boundary.
†Yandangornis			A genus of basal long-tailed avialan. May potentially be a non-avialan genus of theropod.

Mammals

Mammals of the Campanian
Taxa	Presence	Location	Description	Images
Alphadon			A basal metatherian.
Didelphodon			A basal metatherian.
Kamptobaatar			A multituberculate allotherian.
Kennalestes			An insectivorous leptictid eutherian.
Kryptobaatar			Also known as Gobibaatar or Tugrigbaatar. It's a multituberculate allotherian.
Zalambdalestes			Type genus of Zalambdalestidae, a basal clade of Asian eutherians.