2023 in paleoichthyology
| |||
|---|---|---|---|
This list of fossil fish research presented in 2023 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes, and other fishes that were described during the year, as well as other significant discoveries and events related to paleoichthyology that occurred in 2023.
Jawless vertebrates[edit]
| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Sp. nov |
Valid |
Blom et al. |
Devonian (Emsian) |
Shevchenkinskaya Formation |
|
A member of Thelodonti belonging to the group Thelodontiformes and the family Turiniidae. |
||
|
Gen. et sp. nov |
Huang |
Early Cretaceous |
|
A lamprey. The type species is C. miraculum. |
| |||
|
Sp. nov |
Valid |
Zhang et al. |
Silurian (Telychian) |
|
A member of Galeaspida belonging to the family Dayongaspidae. |
|||
|
Gen. et sp. nov |
Gai et al. |
Devonian (Pragian) |
Xiaoshan Formation |
|
A member of Galeaspida belonging to the group Polybranchiaspidiformes and the family Duyunolepididae. The type species is F. novemura. |
| ||
|
Sp. nov |
Liu et al. |
Silurian (Telychian) |
Tataertag Formation |
|
A member of Galeaspida belonging to the family Shuyuidae. |
|||
|
Gen. et sp. nov |
Valid |
McCoy et al. |
Carboniferous (Pennsylvanian) |
A member of Cyclostomi. The type species is S. testai. |
||||
|
Nom. et sp. nov |
Shan et al. |
Silurian |
Tataertag Formation |
|
A member of Galeaspida belonging to the family Xiushuiaspidae; a replacement name for Xiyuaspis Liu et al. (2019). Shan et al. (2023) also named a new species X. lixiensis from the Telychian Qingshui Formation (Jiangxi, China). |
|||
|
Gen. 2 sp. nov |
Wu, Janvier & Zhang |
|
A lamprey. The type species is Y. occisor; genus also includes Y. ingensdentes. |
|
Jawless vertebrate research[edit]
- A study on the anatomy and affinities of Lasanius is published by Reeves et al. (2023), who interpret this vertebrate as a stem-cyclostome.[9]
- Dearden et al. (2023) describe the cranial anatomy of Eriptychius americanus, provide evidence of the presence of a symmetrical set of cartilages interpreted as the preorbital neurocranium, and report that the studied cartilages filled out the head and closely supported the dermal skeleton (in that they were closer to the cranial anatomy of osteostracans and galeaspids than cyclostomes), but were not fused into a single unit around the brain (more closely resembling the cranial anatomy of cyclostomes than osteostracans, galeaspids and jawed vertebrates in that aspect).[10]
- A study on the interaction of fluid flow with 2D models of heterostracan oral plate denticles is published by Grohganz et al. (2023), who interpret their findings as indicating that the studied denticles were not an adaptation to suspension feeding.[11]
Placoderms[edit]
| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Sp. nov |
Valid |
Dupret et al. |
Devonian (Givetian) |
Valentia Slate Formation |
|
|||
|
Gen. et sp. nov |
Valid |
Liu et al. |
Devonian |
Tanglishu Formation |
|
A member of the family Bothriolepididae. The type species is C. magniporus. |
||
|
Sp. nov |
Lebedev in Lebedev et al. |
Devonian (Famennian) |
Bilovo Formation |
|
||||
|
Sp. nov |
Valid |
Plax & Lukševičs |
Devonian (probably Emsian) |
Lepel Beds |
|
A member of Asterolepidoidei belonging to the family Pterichthyodidae. |
||
|
Gen. et sp. nov |
Valid |
Plax & Newman |
Devonian (Emsian) |
|
A member of the family Arctolepididae. The type species is V. profundus. |
Placoderm research[edit]
- Evidence from the study of the skull of Kolymaspis sibirica, interpreted as indicating that the sixth branchial arch was probably the one that was incorporated into the vertebrate shoulder girdle, is presented by Brazeau et al. (2023).[17]
- Brazeau et al. (2023) describe a near-complete "acanthothoracid" upper jaw from the Devonian (Pragian) Yamaat Gol locality (Mongolia), and interpret this finding as indicating that the morphology and function of "acanthoracid" jaws resemble generalized "placoderm" conditions seen also in arthrodires and rhenanids.[18]
- Redescription and a study on the affinities of Bothriolepis sinensis is published by Luo et al. (2023).[19]
- Evidence of different patterns of phylogenetic and taxic diversity of Arthrodira throughout their evolutionary history is presented by Xue et al. (2023), who find evidence robust correlation between declines of phylogenetic diversity and significant global events during the Devonian, especially the late Givetian event, the Late Devonian extinction and the Hangenberg event.[20]
- Engelman (2023) attempts to determine body size of Dunkleosteus terrelli, recovering the body lengths of between 3.1 and 3.5 m for typical adults and ~4.1 m for the largest individuals;[21] in a subsequent study the author reevaluates the methodology and length estimates used by Ferrón, Martinez-Perez & Botella (2017),[22] and argues that length estimates for Dunkleosteus based on the mouth dimensions of extant sharks are not reliable, as arthrodires have proportionally larger mouths than sharks.[23]
- Cui et al. (2023) describe a near-complete post-thoracic exoskeleton of Entelognathus primordialis from the Silurian Kuanti Formation (China), reporting the presence of an anal fin spine in the studied specimen, previously known only in stem cartilaginous fishes, as well as striking similarities of the scales and squamation of the studied specimen to those of bony fishes, including the presence of rhomboid scales with the peg-and-socket articulation previously considered a synapomorphy of bony fishes.[24]
Acanthodians[edit]
| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Gen. et sp. nov |
Valid |
Burrow in Burrow, Murphy & Turner |
Silurian (Přidolí) |
|
An acanthodian of uncertain affinities. The type species is F. nevadaensis. |
|||
|
Gen. et comb. nov |
Valid |
Newman et al. |
Devonian |
Orcadian Basin |
|
A member of Acanthodiformes belonging to the family Mesacanthidae. The type species is "Acanthodes" pusillus Agassiz (1844). |
Acanthodian research[edit]
- Burrow & Desbiens (2023) describe dental elements of Doliodus latispinosus from the Devonian York River Formation (Quebec, Canada), finding no justification for assigning the studied isolated dental elements to a species distinct from D. latispinosus from the Atholville beds (New Brunswick, Canada).[27]
Cartilaginous fishes[edit]
| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Gen. et sp. nov |
Guinot et al. |
Late Cretaceous |
|
A batomorph elasmobranch. The type species is A. acrodonta. |
||||
|
Gen. et sp. nov |
Valid |
Itano |
Carboniferous (late Viséan–early Serpukhovian) |
|
A member of Petalodontiformes belonging to the family Janassidae. The type species is C. whitei. |
|||
|
Sp. nov |
Valid |
Batchelor & Duffin |
Early Cretaceous (Aptian) |
|
A neoselachian shark. Possibly a Carcharhiniforme. |
|||
|
Sp. nov |
Guinot et al. |
Late Cretaceous |
|
A batomorph elasmobranch. |
||||
|
Gen. et sp. nov |
Valid |
Pauliv et al. |
Permian (Asselian) |
Campo Mourão Formation |
|
Possibly a member of the family Symmoriidae. The type species is C. wittigi. |
||
|
Sp. nov |
Guinot et al. |
Late Cretaceous |
|
A batomorph elasmobranch. |
||||
|
Gen. et sp. nov |
Valid |
Marramà et al. |
Eocene (Ypresian) |
|
A member of Myliobatiformes belonging to the new family Dasyomyliobatidae. The type species is D. thomyorkei. |
| ||
|
Sp. nov |
Ivanov in Ivanov, Alekseev & Nikolaeva |
Carboniferous (Viséan) |
|
A member of Symmoriiformes. |
||||
|
Gen. et sp. nov |
Valid |
Ivanov in Ivanov et al. |
Permian |
|
A member of the family Sphenacanthidae. The type species is D. radiata. Published online in 2023, but the issue date is listed as December 2022.[34] |
|||
|
Gen. et sp. nov |
Chahud |
Permian (Cisuralian) |
|
A member of Holocephali. The type species is F. rioclarensis. |
||||
|
Sp. nov |
Tomita, Yabumoto & Kuga |
Oligocene |
Yamaga Formation |
A species of Hemipristis. |
||||
|
Gen. et comb. nov |
Valid |
Lebedev & Itano in Lebedev et al. |
Carboniferous |
|
A member of the family Helicoprionidae. The type species is "Helicoprion" ivanovi Karpinsky (1924). |
|||
|
Sp. nov |
Valid |
Duffin, Heckert & Hancox |
Early Triassic |
|
A member of Hybodontoidea. |
|||
|
Gen. et sp. nov |
Wen et al. |
Middle Triassic (Anisian) |
|
A member of the family Lonchidiidae. Genus includes new species L. striata. |
||||
| Maghriboselache[40] | Gen. et sp. nov | Klug et al. | Late Devonian |  Morocco
|
A member of the family Cladoselachidae. The type species is M. mohamezanei. | 
| ||
|
Sp. nov |
Guinot et al. |
Late Cretaceous |
|
A batomorph elasmobranch. |
||||
|
Sp. nov |
Boulemia & Adnet |
Paleogene |
Kef Esnoun Formation |
|
||||
|
Sp. nov |
Begat et al. |
Late Cretaceous (Cenomanian) |
|
|||||
|
Ptychotrygon nazeensis[28] |
Sp. nov |
Guinot et al. |
Late Cretaceous |
|
A batomorph elasmobranch. |
|||
|
Sp. nov |
Guinot et al. |
Late Cretaceous |
|
A batomorph elasmobranch. |
||||
|
Gen. et sp. nov |
Guinot et al. |
Late Cretaceous |
|
A batomorph elasmobranch. The type species is S. minimus. |
||||
|
Sp. nov |
Ivanov in Ivanov, Alekseev & Nikolaeva |
Carboniferous (Viséan) |
|
|||||
|
Gen. et sp. nov |
Valid |
Hodnett et al. |
Carboniferous (Viséan) |
|
A member of Petalodontiformes belonging to the family Janassidae. The type species is S. tollesonae. |
|||
|
Sp. nov |
Valid |
Stumpf et al. |
Middle Jurassic (Bajocian) |
Agoudim Formation |
|
A member of Hybodontiformes belonging to the family Hybodontidae and the subfamily Acrodontinae. |
| |
|
Gen. et sp. nov |
Valid |
Itano & Duffin |
Carboniferous (Viséan) |
St. Louis Formation |
|
A chimaera belonging to the group Squalorajoidei. The type species is S. schachti. |
||
|
Gen. et sp. nov |
Guinot et al. |
Late Cretaceous |
|
A batomorph elasmobranch. The type species is T. thiami. |
Cartilaginous fish research[edit]
- Fossil material of members of the genera Pucapampella and Zamponiopteron is described from the Devonian (Eifelian) Chagrapi Formation by Zevallos-Valdivia et al. (2023), representing the first record of Paleozoic vertebrates from Peru reported to date.[46]
- A study on the musculoskeletal anatomy of Iniopera is published by Dearden, Herrel & Pradel (2023), who interpret the anatomy of Iniopera as unsuited to durophagy, and consider it to be likely a high-performance suction-feeder.[47]
- Fossil material of members of at least seven species belonging to the genus Ptychodus is described from the ?Cenomanian–Santonian of the Malyy Prolom area (Ryazan Oblast, Russia) by Amadori et al. (2023), who also report the northernmost occurrence of Ptychodus in Europe from the Cenomanian of Varavinsky ravine area (Moscow Oblast, Russia), and interpret the studied fossils as indicating that Late Cretaceous epicontinental seas of the Russian platform were important areas of diversification and spread of Ptychodus.[48]
- Amadori et al. (2023) report the discovery of teeth of various species belonging to the genus Ptychodus from the Cenomanian and Turonian deposits of Ukraine, including teeth of cuspidate (P. altior) and un-cuspidate species (P. decurrens, P. latissimus, P. marginalis and P. polygyrus), and argue that the availability of diverse shelled invertebrates in epicontinental seas might have favored the diversification of Ptychodus.[49]
- Ghosh et al. (2023) report the discovery of a new assemblage of lamniform shark teeth from the Aptian Habur Formation (India), including teeth of Dwardius and possibly of Eostriatolamia which may be some of the globally oldest record of these taxa.[50]
- A study on the teeth of Megachasma applegatei is published by Krak & Shimada (2023), who find that the range of the morphometric variation of teeth of M. applegatei is larger than that of teeth of extant megamouth shark, with different tooth types corresponding to tooth types present in the smalltooth sand tiger.[51]
- Shimada et al. (2023) describe tessellated calcified cartilage and placoid scale associated with a tooth set of Otodus megalodon from the Miocene strata in Japan, and interpret the morphology of the studied material as indicating that O. megalodon was generally a slow cruising shark.[52]
- A study on the thermoregulation in Otodus megalodon is published by Griffiths et al. (2023), who argue that O. megalodon had an overall warmer body temperature compared with other coexisting shark species, and that its large body size coupled with high metabolic costs associated with having at least partial endothermy might have made it vulnerable to extinction.[53]
- Collareta, Casati & Di Cencio (2023) describe new fossil material of Parotodus benedenii from the Valdelsa Basin (Italy), providing evidence of the survival of the species at least until the Late Pliocene, and interpret P. benedenii as a large-bodied carnivorous shark living in pelagic settings.[54]
- Collareta et al. (2023) report the discovery of teeth of Alopias grandis from the Miocene deposits in southern Italy, possibly including the geologically youngest record of the species and extending its known geographic range.[55]
- Villafaña et al. (2023) describe fossil material of the common thresher and the porbeagle from the Bahía Inglesa Formation (Caldera Basin, Chile), confirming the abundance of lamniform sharks in the Eastern Pacific of South America during the Neogene.[56]
- Ehret et al. (2023) provisionally refer the species Cosmopolitodus planus/Isurus planus to the genus Carcharodon, and describe fossil material of C. planus and Carcharodon hubbelli from Miocene deposits in the South Island, representing the first records of both species from New Zealand reported to date.[57]
- A study on the anatomy and affinities of Protospinax annectans, based on data from both known and previously undescribed specimens from the Tithonian Altmühltal Formation (Germany), is published by Jambura et al. (2023).[58]
- Ferrón (2023) argues that, although representatives of most squalomorph groups colonized deep waters independently during the Late Jurassic and Early Cretaceous, bioluminescence evolved only once among sharks in a bathydemersal ancestor.[59]
- A fossil egg case containing a well-preserved batoid (possibly stem-myliobatiform) embryo, with a unique combination of characters indicating that the embryo represents a previously unknown batoid form, is described from the Cenomanian Sannine limestone of Hjoula (Lebanon) by Capasso & Yamaguchi (2023).[60]
- Reinecke et al. (2023) study the anatomy and affinities of whiptail stingray teeth from the Chattian of northern Germany and the Burdigalian of southern France, transferring the species Dasyatis probsti to the genus Bathytoshia.[61]
- Pollerspöck et al. (2023) describe an assemblage of deep-sea shark fossils from the Eocene (Ypresian) Lillebælt Clay Formation (Denmark), showing highest similarities with deep-sea shark faunas of France, Austria and northern Morocco in spite of the North Sea Basin having lost direct connections to the neighbouring marine areas in the Eocene.[62]
- Kovalchuk et al. (2023) revise the taxonomic composition of the cartilaginous fish assemblage from the Eocene (Lutetian-Bartonian) Kyiv Formation (Ukraine), interpreting the studied taxa as inhabiting shallow, warm waters and confined to the continental shelf.[63]
- Verma (2023) describes new fossil material of elasmobranchs from the Eocene (Bartonian) Harudi Formation (India), providing evidence of replacement of earlier Eocene assemblages of elasmobranchs from western India by an assemblage dominated by members of the genera Brachycarcharias, Striatolamia, Galeocerdo and Carcharhinus, which might have been linked to the Middle Eocene Climatic Optimum.[64]
- An assemblage of shark and ray teeth, interpreted as indicative of a warm, shallow water community, is described from the Lower Miocene deposits of the Upper Marine Molasse near Ballendorf (Germany) by Höltke et al. (2023).[65]
- A study on changes of diversity of European chondrichthyans during the Neogene is published by Villafaña et al. (2023).[66]
- A study on the impact of the Cretaceous–Paleogene extinction event on elasmobranchs is published by Guinot & Condamine (2023), who find rays and durophagous species to be more affected by the extinction than sharks and nondurophagous species, and find taxa with large geographic ranges or restricted to high-latitude settings to show higher survival.[67]
Ray-finned fishes[edit]
| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A species of Amblyeleotris. |
|||
|
Gen. et sp. nov |
Valid |
Štamberg & Werneburg |
Permian (Asselian) |
Lower Goldlauter Formation |
|
A member of the family Aeduellidae. The type species is A. ornata. |
||
|
Sp. nov |
Pindakiewicz, Hryniewicz & Kaim |
Early Cretaceous (Valanginian) |
|
|||||
|
Sp. nov |
Valid |
Bannikov & Erebakan |
Oligocene |
|
A member of the family Carangidae. |
|||
|
Sp. nov |
Valid |
Marramà & Carnevale |
Early Cretaceous (Albian) |
|
A member of Clupeomorpha belonging to the group Ellimmichthyiformes and the family Armigatidae. |
|||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A scaldfish. |
|||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A scaldfish. |
|||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A species of Aseraggodes. |
|||
|
Sp. nov |
Lin in Lin et al. |
Pliocene |
Gutingkeng Formation |
Taiwan |
A species of Benthosema. |
|||
|
Sp. nov |
Lin in Lin et al. |
Pliocene |
Gutingkeng Formation |
Taiwan |
A species of Benthosema. |
|||
|
Gen. et sp. nov |
Murray, Nelson & Brinkman |
Late Cretaceous (Campanian) |
|
A sturgeon. The type species is B. labyrinthicus. |
||||
|
Sp. nov |
Valid |
Schwarzhans & von der Hocht |
Miocene |
A species of Bothus. |
||||
|
Sp. nov |
Valid |
Schwarzhans & Nielsen |
Eocene |
Lillebælt Clay Formation |
|
A codlet. |
||
|
Gen. et sp. nov |
Schrøder et al. |
Eocene |
|
A stromateoiform, possibly a medusafish. The type species is B. henricii. |
||||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A species of Callionymus. |
|||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A species of Callionymus. |
|||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A species of Capros. |
|||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Caranx. |
||||
|
Sp. nov |
Valid |
Schwarzhans & von der Hocht |
Miocene |
A species of Cataetyx. |
||||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Centroberyx. |
||||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Cepola. |
||||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A species of Chelon. |
|||
|
Gen. et sp. nov |
Valid |
De Mayrinck et al. |
Early Cretaceous (Aptian) |
Barbalha Formation |
|
A member of the family Cladocyclidae. The type species is C. araripensis. |
||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
Possibly a species of Congiopodus. |
|||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Deltentosteus. |
||||
|
Sp. nov |
Chida, Brinkman & Murray |
Late Cretaceous (Campanian) |
|
A member of the family Dercetidae. |
||||
|
Gen. et sp. nov |
Xu & Ma |
Middle Triassic (Anisian) |
|
A basal member of Ginglymodi. The type species is D. ocellatus. |
||||
|
Sp. nov |
Valid |
Schwarzhans & Nielsen |
Eocene |
Lillebælt Clay Formation |
|
Possibly a species of Diaphus. |
||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Diaphus. |
||||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A species of Dicologlossa. |
|||
|
Sp. nov |
Valid |
Schwarzhans et al. |
Early Miocene |
|
A species of Galaxias. |
|||
|
Sp. nov |
Valid |
Schwarzhans et al. |
Early Miocene |
Bannockburn Formation |
|
A species of Galaxias. |
||
|
Sp. nov |
Valid |
Schwarzhans et al. |
Early Miocene |
Bannockburn Formation |
|
A species of Galaxias. |
||
|
Sp. nov |
Valid |
Schwarzhans et al. |
Early Miocene |
Bannockburn Formation |
|
A species of Galaxias. |
||
|
Sp. nov |
Valid |
Schwarzhans et al. |
Early Miocene |
Bannockburn Formation |
|
A species of Galaxias. |
||
|
Gen. et 2 sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A goby belonging to the subfamily Gobiinae and to the Aphia lineage. The type species is G. globulosus; genus also includes G. depressus. |
|||
|
Gen. et sp. nov |
Cooper et al. |
Late Cretaceous (Turonian) |
Akrabou Formation |
|
A gar. The type species is G. gherisensis. |
| ||
|
Gen. et sp. nov |
Lin in Lin et al. |
Pliocene |
Gutingkeng Formation |
Taiwan |
A viviparous brotula. The type species is G. changi. |
|||
|
Sp. nov |
Valid |
Schwarzhans & von der Hocht |
Miocene |
A species of Hygophum. |
||||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A species of Hyrcanogobius. |
|||
|
Gen. et sp. nov |
Valid |
Andrews et al. |
|
A stem-lineage member of Holocentridae. |
||||
|
Gen. et sp. nov |
Valid |
Deesri et al. |
Early Cretaceous (Aptian) |
|
A member of the family Amiidae belonging to the subfamily Sinamiinae. The type species is K. phattharajani. |
|||
|
Sp. nov |
Pindakiewicz, Hryniewicz & Kaim |
Early Cretaceous (Valanginian) |
|
|||||
|
Gen et 2 sp. nov |
Valid |
Bakaev in Esin & Bakaev |
Permian |
|
An early ray-finned fish belonging to the group Palaeonisciformes and the family Palaeoniscidae. The type species is K. gubini Esin & Bakaev; genus also includes K. dozmerensis Esin & Bakaev. Published online in 2023, but the issue date is listed as December 2022.[86] |
|||
|
Gen. et sp. nov |
Valid |
Recinos et al. |
Late Cretaceous (Campanian) |
|
A bonefish. The type species is M. tojolabalensis. |
|||
|
Gen. et comb. et sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
|
A member of the family Clupeidae belonging to the subfamily Alosinae. The type species is "Otolithus (Osmeridarum)" wilhelmi Djafarova (2006); genus also includes "Clupea" gomotartziensis Strashimirov (1985), as well as new species Maeotichthys salebrosus Schwarzhans & Bratishko in Schwarzhans, Bratishko & Vernyhorova (2023).[88] |
|||
|
Sp. nov |
Valid |
Schwarzhans et al. |
Early Miocene |
Bannockburn Formation |
|
A species of Mataichthys. |
||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A species of Mesogobius. |
|||
|
Gen et sp. nov |
Valid |
Bulanov, Minikh & Golubev |
Permian |
|
A member of Eurynotoidiformes. The type species is M. primus. Published online in 2023, but the issue date is listed as December 2022.[89] |
|||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Myripristis. |
||||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A species of Neogobius. |
|||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A species of Neogobius. |
|||
|
Sp. nov |
Valid |
Capasso |
Late Cretaceous (Turonian) |
Akrabou Formation |
A member of Pycnodontiformes belonging to the family Pycnodontidae. |
|||
|
Sp. nov |
In press |
Yabumoto & Zhang |
Miocene |
Chojabaru Formation |
|
A species of Odontobutis. |
||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Ophidion. |
||||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A member of the family Sparidae. |
||||
|
Sp. nov |
Valid |
Schwarzhans & von der Hocht |
Miocene |
|||||
|
Gen. et sp. nov |
Pindakiewicz, Hryniewicz & Kaim |
Early Cretaceous (Valanginian) |
|
Genus includes new species P. plicata. |
||||
|
Sp. nov |
Valid |
Stringer & Sloan |
Late Cretaceous (Maastrichtian) |
|
A member of the family Merlucciidae. |
|||
|
Gen. et sp. nov |
Valid |
Caballero-Viñas, Alvarado-Ortega & Cantalice Severiano |
Pliocene |
|
A member of the family Goodeidae belonging to the subfamily Goodeinae. The type species is P. guzmanae. |
|||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A member of the family Gadidae. |
|||
|
Sp. nov |
Valid |
Marramà & Carnevale |
Early Cretaceous (Albian) |
Pietraroja Plattenkalk |
|
A member of Clupeomorpha belonging to the group Ellimmichthyiformes and the family Paraclupeidae. |
||
|
Gen. et sp. et comb. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
|
A member of the family Macroramphosidae. The type species is P. pumilis; genus also includes "Оtolithus (inc. sedis)" platessaeformis Pobedina (1956). |
|||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Parapristipoma. |
||||
|
Gen. et sp. nov |
Valid |
Hilton et al. |
Late Cretaceous (Maastrichtian) |
|
A paddlefish. The type species is P. willybemisi. |
| ||
|
Sp. nov |
Valid |
Schwarzhans & von der Hocht |
Miocene |
A species of Parascolopsis. |
||||
|
Gen. et sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A dartfish. The type species is P. alienus. |
|||
|
Gen. et sp. nov |
Valid |
Caron et al. |
Carboniferous (Bashkirian) |
Marsden Formation |
|
An early ray-finned fish, possibly stem-neopterygian. The type species is P. mossae. |
||
|
Sp. nov |
Brito & Vullo |
Late Cretaceous (Cenomanian) |
|
|||||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Pomadasys. |
||||
|
Gen. et 3 sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A goby belonging to the subfamily Gobiinae and to the Benthophilus lineage. The type species is P. ahnelti; genus also includes P. trigonus and P. zonatus. |
|||
|
Gen. et sp. et comb. nov |
Valid |
Schwarzhans & Nielsen |
Eocene |
Lillebælt Clay Formation |
|
A cusk-eel. The type species is P. schnetleri; genus also includes "Neobythites" bozzolo Lin, Nolf & Girone in Lin et al. (2016) and "Neobythites" leonardi Lin, Nolf & Girone in Lin et al. (2016). |
||
|
Sp. nov |
Pindakiewicz, Hryniewicz & Kaim |
Early Cretaceous (Valanginian) |
|
|||||
|
Sp. nov |
Pindakiewicz, Hryniewicz & Kaim |
Early Cretaceous (Valanginian) |
|
|||||
|
Sp. nov |
Pindakiewicz, Hryniewicz & Kaim |
Early Cretaceous (Valanginian) |
|
|||||
|
Gen. et sp. nov |
Valid |
Hilton et al. |
Late Cretaceous (Maastrichtian) |
Hell Creek Formation |
|
A paddlefish. The type species is P. inundatus. |
||
|
Gen. et sp. nov |
Valid |
El Hossny et al. |
Late Cretaceous (Cenomanian) |
Hesseltal Formation |
A member of the family Plethodidae. The type species is R. taxidiotis. El Hossny et al. (2023) also interpret fossils of "Protosphyraena" minor and "Protosphyraena" stebbingi as fossil material of Rhamphoichthys sp., but don't consider these to be valid taxa due to the incompleteness of their fossil material. |
| ||
|
Sp. nov |
Valid |
Calzoni et al. |
Eocene (Ypresian) |
|
A member of Syngnathiformes belonging to the group Dactylopteroidei and the family Rhamphosidae. |
|||
|
Rhamphosus brevirostris[98] |
Sp. nov |
Valid |
Calzoni et al. |
Eocene (Ypresian) |
Monte Bolca |
|
A member of Syngnathiformes belonging to the group Dactylopteroidei and the family Rhamphosidae. |
|
|
Rhamphosus longispinatus[98] |
Sp. nov |
Valid |
Calzoni et al. |
Eocene (Ypresian) |
Monte Bolca |
|
A member of Syngnathiformes belonging to the group Dactylopteroidei and the family Rhamphosidae. |
|
|
Rhamphosus tubulirostris[98] |
Sp. nov |
Valid |
Calzoni et al. |
Eocene (Ypresian) |
Monte Bolca |
|
A member of Syngnathiformes belonging to the group Dactylopteroidei and the family Rhamphosidae. |
|
|
Sp. nov |
Schwarzhans |
Miocene (Tortonian) and Pliocene (Zanclean) |
|
A species of Rhynchoconger. |
||||
|
Sp. nov |
Valid |
Fang et al. |
|
Announced in 2022; the final article version was published in 2023. |
||||
|
Sp. nov |
Valid |
Carrillo-Briceño et al. |
Miocene |
|
A species of Sciades. |
|||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A species of Scomber. |
|||
|
Gen. et sp. nov |
Valid |
Schwarzhans & Bratishko in Schwarzhans, Bratishko & Vernyhorova |
Miocene |
Crimea |
A member of the family Gobiidae belonging to the tribe Benthophilini. The type species is S. spissus. |
|||
|
Sp. nov |
Valid |
Nazarkin |
Miocene |
Kurasi Formation |
|
A species of Serrivomer. |
||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Spondyliosoma. |
||||
|
Gen. et sp. nov |
Valid |
Capasso |
Late Cretaceous (Cenomanian) |
|
A member of Pycnodontiformes. The type species is S. elongata. |
|||
|
Gen. et sp. nov |
Valid |
Schrøder & Carnevale |
Eocene (Ypresian) |
|
A member of Argentiniformes. The type is species S. longigracilis. |
|||
|
Sp. nov |
Than-Marchese et al. |
Late Cretaceous (Cenomanian) |
Cintalapa Formation |
|
A member of Clupeomorpha belonging to the group Ellimmichthyiformes. |
|||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Trachinus. |
||||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Trachinus. |
||||
|
Sp. nov |
Valid |
Schwarzhans & von der Hocht |
Miocene |
A species of Trachurus. |
||||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Trachurus. |
||||
|
Sp. nov |
Valid |
Hoedemakers in De Schutter et al. |
Oligocene (Rupelian) |
Boom Formation |
|
A species of Trachurus. |
||
|
Gen. et comb. nov |
Valid |
Schwarzhans & Nielsen |
Eocene |
Lillebælt Clay Formation |
|
A member of Acanthopterygii of uncertain affinities. The type species is "Caproidarum" madseni Schwarzhans (2007). |
||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Uranoscopus. |
||||
|
Sp. nov |
Schwarzhans |
Pliocene (Zanclean) |
|
A species of Uranoscopus. |
||||
|
Gen. et sp. nov. |
Murray et al. |
Late Cretaceous (Maastrichtian) |
Mahajanga Basin |
|
A member of the family Chanidae. The type species is V. fahiny. |
|||
|
Gen et comb. nov |
Valid |
Bulanov, Minikh & Golubev |
Permian |
Poldarsa/Poldarskaya Formation |
|
A member of Eurynotoidiformes. The type species is "Isadia" opokiensis Minikh & Andrushkevich (2017). Published online in 2023, but the issue date is listed as December 2022.[89] |
||
|
Gen. et sp. nov |
Valid |
El Hossny & Cavin |
Late Cretaceous (Cenomanian) |
|
A teleost of uncertain affinities. The type species is Y. sibelleae. |
| ||
|
Sp. nov |
Valid |
Bratishko & Schwarzhans in Bratishko, Schwarzhans & Vernyhorova |
Miocene |
Crimea |
A species of Zosterisessor. |
Ray-finned fish research[edit]
- Figueroa et al. (2023) report brain and cranial nerve soft-tissue preservation in the type specimen of Coccocephalus wildi from the Carboniferous strata in the Mountain Fourfoot Mine (Pennine Lower Coal Measures; Lancashire, United Kingdom).[108]
- Bakaev, Johanson & LeBlanc (2023) study the dental system of Kazanichthys viatkensis, reporting the presence of morphological similarities to the dental systems of extant sparids, and interpreting K. viatkensis as a generalist durophagous feeder.[109]
- Revision of the fossil material of Permian ray-finned fishes from the Kazankovo-Markino Formation (Kemerovo Oblast, Russia) is published by Bakaev (2023), who considers Heterolepis Sergienko (1974) to be a junior synonym of Eurynotoides Berg (1940).[110]
- Martill (2023) describes a bony scute of a sturgeon from the Maastrichtian marine phosphatites of central Morocco, representing the first record of an acipenseriform fish from Africa reported to date.[111]
- New information on the morphology of the scales of members of the family Pseudobeaconiidae, based on new fossil material from the Triassic Santa Clara Abajo Formation (Argentina), is presented by Giordano, Benavente & Suárez (2023).[112]
- Putative eugnathid amiiform Sinoeugnathus kueichowensis is reinterpreted as a small-sized member of Ionoscopiformes by Feng et al. (2023), who name a new family Subortichthyidae including the genera Subortichthys, Sinoeugnathus, Allolepidotus and Eoeugnathus.[113]
- Sullivan, Jasinski & Williamson (2023) describe an exceptionally well-preserved articulated skull roof and braincase of Melvius chauliodous from the Upper Cretaceous Kirtland Formation (New Mexico, United States), revise the characters that defin the genus and its two recognized species, and study the phylogenetic affinities of Melvius.[114]
- A study on the microstructure of teeth of Late Jurassic pachycormids and caturoids from the Owadów-Brzezinki site (Poland) is published by Weryński, Błażejowski & Kędzierski (2023), who report structural differences interpreted as suggestive of different adaptations for predation and possible niche partitioning between the studied taxa.[115]
- Systematic revision of the Late Jurassic species of Caturidae is published by López-Arbarello & Ebert (2023).[116]
- Fossil material of a putative member of the genus Caturus reported by Bogan, Taverne & Agnolin (2013) as found in the Triassic Los Menucos Group,[117] is reinterpreted by López-Arbarello et al. (2023) as actually collected in outcrops of the Jurassic Vaca Muerta Formation, and excluded from the genus Caturus.[118]
- Cooper & Maxwell (2023) describe a specimen of Pachycormus macropterus from the Toarcian Posidonia Shale (Germany) preserved with an unusually large ammonite inside its gut, interpreted as ingested immediately prior to and directly responsible for the fish's death.[119]
- A study on the bone histology of Araripichthys castilhoi, interpreted as corroborating its placement within basal Teleostei, is published by Mayrinck et al. (2023).[120]
- Stinnesbeck et al. (2023) report the presence of two different body shape types of specimens of Tselfatia formosa from the Turonian platy limestone deposit of Vallecillo (Mexico), intepreted as evidence of sexual dimorphism, and interpret the anatomy of its fins as indicating that T. formosa lived in a deep water environment and that its lifestyle resembled that of extant fan fishes.[121]
- Cooper & Norton (2023) describe fossil material of an indeterminate plethodid from the Maastrichtian deposits from the Plateau des Phosphates (Morocco), representing the youngest occurrence of a plethodid reported to date.[122]
- Redescription and a study on the affinities of Sorbinichthys elusivo is published by Taverne & Capasso (2023).[123]
- Fossil material of a catfish, representing the first record of a bony fish from the Maastrichtian of the Marília Formation (Brazil) and extending known Late Cretaceous catfish distribution, is described by Candeiro et al. (2023).[124]
- A study on the fossil record of acanthomorphs from the Maastrichtian–Paleocene strata is published by Friedman et al. (2023), who find that the majority of the principal acanthomorph groups appear in the fossil record before the end of the Paleocene.[125]
- A study on the variety of the morphology of the first abdominal vertebral centrum in extant acanthomorphs is published by Murray & Brinkman (2023), who interpret their findings as indicating that the overall morphology of the first centrum is conservative within acanthomorph families, and that it is possible to assign many fossil acanthomorph centra to extant families, suborders or orders.[126]
- Rust & Robinson (2023) redescribe Eothyrsites holosquamatus, and interpret this taxon as likely representing an ancestral form of gempylid.[127]
- Fossil material representing one of the oldest records of marlins reported to date is described from the Miocene (Aquitanian) Northern Alpine Foreland Basin (Austria) by De Gracia, Berning & Kriwet (2023), who report evidence of coexistence of marlins, xiphiorhynchine xiphiids and aglyptorhynchine palaeorhynchids from the Northern Alpine Foreland Basin and from the Oligocene Chandler Bridge Formation (South Carolina, United States).[128]
- Ngoepe et al. (2023) reconstruct the history of arrival order and relative abundances of major fish groups from Lake Victoria, using data from the continuous fossil record from the preceding 17,000 years, and report that cichlids did not dominate the assemblage until several thousand years into its history, but they were the only major group that had the ecological versatility that allowed them to persist once the new deep and open-water habitats emerged.[129]
- Evidence from (mostly lanternfish) otoliths from the Lindos Bay Formation (Rhodes, Greece), interpreted as indicative of an overall decline of the median size of lanternfishes in the eastern Mediterranean during MIS 19 interglacial, but also as indicative of different trends in size in individual mesopelagic species across the studied time interval, is presented by Agiadi et al. (2023).[130]
Lobe-finned fishes[edit]
| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Gen. et sp. nov |
Valid |
Elliott, Challands & Smithson |
Carboniferous |
|
A lungfish. The type species is B. kerri. |
|||
|
Sp. nov |
Valid |
Minikh |
Triassic |
|
A lungfish. Published online in 2023, but the issue date is listed as December 2022.[132] |
|||
|
Sp. nov |
Downs et al. |
Devonian (Famennian) |
|
|||||
|
Sp. nov |
Valid |
Gess & Ahlberg |
Devonian (Famennian) |
|
| |||
|
Gen. et sp. nov |
Valid |
Johanson et al. |
Devonian (Pragian) |
|
A lungfish. The type species is J. tuulingi. |
|||
|
Gen. et sp. nov |
Valid |
Elliott, Challands & Smithson |
Carboniferous |
|
A lungfish. The type species is L. clarki. |
|||
|
Sp. nov |
Elliott |
Carboniferous |
|
A member of the family Megalichthyidae. |
||||
|
Gen. et sp. nov |
Valid |
Ferrante & Cavin |
Middle Triassic (Anisian) |
|
A coelacanth belonging to the family Latimeriidae. The type species is R. heinzfurreri. |
| ||
|
Sp. nov |
Valid |
Ferrante et al. |
Middle Triassic |
Prosanto Formation |
|
A coelacanth belonging to the family Latimeriidae. |
||
| Whiteia giganteus[139] | Sp. nov | Brownstein | Late Triassic | Dockum Group |  United States ( Texas)
|
A coelacanth. |
Lobe-finned fish research[edit]
- Dupret et al. (2023) describe new fossil material of sarcopterygians from the Devonian (Givetian) Valentia Slate Formation (Republic of Ireland), including a tooth plate of a lungfish with a derived morphology otherwise only known from Late Devonian and later taxa, and a possible rhizodontid fossil material, which might indicate that a dispersal of rhizodontids from Gondwana into Euramerica happened as early as middle Givetian.[140]
General research[edit]
- Kuznetsov & Kryukova (2023) present new reconstructions of subcephalic musculature for Pucapampella, Eusthenopteron and Ichthyostega.[141]
- Baucon et al. (2023) describe fish-feeding traces from the Lower Cretaceous (Hauterivian–Barremian) Palombini Shale Formation (Italy), interpreted as the earliest direct evidence of bottom-living vertebrates from the deep sea.[142]
- Trif et al. (2023) describe a diverse fish assemblage from the Priabonian strata in the Leghia-Tabără area (Romania), including the first record of Physogaleus alabamensis from Europe and the first record of Striatolamia tchelkarnurensis outside the Turgai Strait region.[143]
References[edit]
- ^ Blom, H.; Vaškaninová, V.; Karatajūtė-Talimaa, V.; Žigaitė, Ž. (2023). "Thelodont scales from the Lower Devonian of Novaya Zemlya Archipelago, Arctic Russia". Spanish Journal of Palaeontology. 38 (1): 9–14. doi:10.7203/sjp.26231. S2CID 257676099.
- ^ Huang, W. (2023). "A new species of fossil lamprey (Petromyzontida: Petromyzontiformes) from Hebei, China". Historical Biology: An International Journal of Paleobiology: 1–13. doi:10.1080/08912963.2023.2252443. S2CID 261473806.
- ^ Zhang, Y.; Li, X.; Shan, X.; Lin, X.; Tan, K.; Li, Q.; Zhao, W.; Tang, L.; Zhu, M.; Gai, Z. (2023). "The first galeaspid fish (stem-Gnathostomata) from the Silurian Xiushan formation of Hunan Province, China". Historical Biology: An International Journal of Paleobiology: 1–12. doi:10.1080/08912963.2023.2225083. S2CID 259469524.
- ^ Gai, Z.; Lin, X.; Shan, X.; Ferrón, H. G.; Donoghue, P. C. J. (2023). "Postcranial disparity of galeaspids and the evolution of swimming speeds in stem-gnathostomes". National Science Review. 10 (7). nwad050. doi:10.1093/nsr/nwad050. PMC 10232041. PMID 37266551.
- ^ Liu, W.; Shan, X.; Lin, X.; Shen, Y.; Liu, Y.; Zhang, Z.; Gai, Z. (2023). "The first Eugaleaspiforme fish from the Silurian of the Tarim Basin reveals a close relationship between the Tarim and South China blocks at 438 mya". Palaeogeography, Palaeoclimatology, Palaeoecology. 628. 111774. Bibcode:2023PPP...62811774L. doi:10.1016/j.palaeo.2023.111774. S2CID 260874354.
- ^ McCoy, V. E.; Wittry, J.; Sadabadi, H.; Mayer, P. (2023). "A reappraisal of Nemavermes mackeei from the Mazon Creek fossil site expands Carboniferous cyclostome diversity". Journal of Paleontology. 97 (5): 1116–1132. Bibcode:2023JPal...97.1116M. doi:10.1017/jpa.2023.72.
- ^ Shan, X.-R.; Lin, X.-H.; Zhang, Y.-M.; Li, X.-T.; Gai, Z.-K. (2023). "New findings of Xiyuichthys (Xiushuiaspidae, Galeaspida) from the Silurian of Jiangxi Province and Tarim Basin". Vertebrata PalAsiatica. 61 (4): 245–260. doi:10.19615/j.cnki.2096-9899.230904.
- ^ Wu, F.; Janvier, P.; Zhang, C. (2023). "The rise of predation in Jurassic lampreys". Nature Communications. 14 (1). 6652. Bibcode:2023NatCo..14.6652W. doi:10.1038/s41467-023-42251-0. PMC 10618186. PMID 37907522.
- ^ Reeves, J. C.; Wogelius, R. A.; Keating, J. N.; Sansom, R. S. (2023). "Lasanius, an exceptionally preserved Silurian jawless fish from Scotland". Palaeontology. 66 (2). e12643. Bibcode:2023Palgy..6612643R. doi:10.1111/pala.12643. S2CID 258066900.
- ^ Dearden, R. P.; Lanzetti, A.; Giles, S.; Johanson, Z.; Jones, A. S.; Lautenschlager, S.; Randle, E.; Sansom, I. J. (2023). "The oldest three-dimensionally preserved vertebrate neurocranium". Nature. 621 (7980): 782–787. Bibcode:2023Natur.621..782D. doi:10.1038/s41586-023-06538-y. PMC 10533405. PMID 37730987.
- ^ Grohganz, M.; Ferrón, H. G.; Johanson, Z.; Donoghue, P. C. J. (2023). "Testing hypotheses of pteraspid heterostracan feeding using computational fluid dynamics". Journal of Vertebrate Paleontology. 43 (2). e2272974. doi:10.1080/02724634.2023.2272974.
- ^ Dupret, V.; Byrne, H. M.; Castro, N.; Hammer, Ø.; Higgs, K. T.; Long, J. A.; Niedźwiedzki, G.; Qvarnström, M.; Stössel, I.; Ahlberg, P. E. (2023). "The Bothriolepis (Placodermi, Antiarcha) material from the Valentia Slate Formation of the Iveragh Peninsula (middle Givetian, Ireland): Morphology, evolutionary and systematic considerations, phylogenetic and palaeogeographic implications". PLOS ONE. 18 (2). e0280208. Bibcode:2023PLoSO..1880208D. doi:10.1371/journal.pone.0280208. PMC 9949654. PMID 36821588.
- ^ Liu, S.; Pan, Z.; Zhu, M.; Jia, L.; Zhao, W. (2023). "A New Bothriolepid Antiarch from the Middle Devonian in Luquan, Yunnan, South China and Its Stratigraphic and Biogeographic Significances". Journal of Earth Science. 34 (4): 1176–1188. Bibcode:2023JEaSc..34.1176L. doi:10.1007/s12583-022-1775-1. S2CID 260813014.
- ^ Lebedev, O. A.; Engelman, R. K.; Skutschas, P. P.; Johanson, Z.; Smith, M. M.; Kolchanov, V. V.; Trinajstic, K.; Linkevich, V. V. (2023). "Structure, Growth and Histology of Gnathal Elements in Dunkleosteus (Arthrodira, Placodermi), with a Description of a New Species from the Famennian (Upper Devonian) of the Tver Region (North-Western Russia)". Diversity. 15 (5). 648. doi:10.3390/d15050648.
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