The evolutionary ancestry of arthropods dates back to the Cambrian period. The group is generally regarded as monophyletic, and many analyses support the placement of arthropods with cycloneuralians (or their constituent clades) in a superphylum Ecdysozoa. Overall, however, the basal relationships of animals are not yet well resolved. Likewise, the relationships between various arthropod groups are still actively debated. Today, arthropods contribute to the human food supply both directly as food, and more importantly, indirectly as pollinators of crops. Some species are known to spread severe disease to humans, livestock, and crops. (Full article...)
Prionomyrmecini is an anttribe belonging to the subfamily Myrmeciinae established by William Morton Wheeler in 1915. Two members are a part of this tribe, the extant Nothomyrmecia and the extinct Prionomyrmex. The tribe was once considered a subfamily due to the similarities between Nothomyrmecia and Prionomyrmex, but such reclassification was not widely accepted by the scientific community. These ants can be identified by their long slender bodies, powerful stingers and elongated mandibles. Fossil Prionomyrmecini ants were once found throughout Europe, possibly nesting in trees and preferring jungle habitats. Today, Prionomyrmecini is only found in Australia, preferring old-growth mallee woodland surrounded by Eucalyptus trees. Nothomyrmecia workers feed on nectar and arthropods, using their compound eyes for prey and navigational purposes. Owing to their primitive nature, they do not recruit others to food sources or create pheromone trails. Nothomyrmecia colonies are small, consisting of 50 to 100 individuals. (Full article...)
The antlions are a group of about 2,000 species of insect in the neuropteranfamilyMyrmeleontidae. They are known for the predatory habits of their larvae, which mostly dig pits to trap passing ants or other prey. In North America, the larvae are sometimes referred to as doodlebugs because of the marks they leave in the sand. The adult insects are less well known due to their relatively short lifespans compared to the larvae. Adults, sometimes known as antlion lacewings, mostly fly at dusk or just after dark and may be mistakenly identified as dragonflies or damselflies.
Antlions have a worldwide distribution. The greatest diversity occurs in the tropics, but a few species are found in cold-temperate locations, one such being the European Euroleon nostras. They most commonly occur in dry and sandy habitats where the larvae can easily excavate their pits, but some larvae hide under debris or ambush their prey among leaf litter.
Antlions are poorly represented in the fossil record. Myrmeleontiformia is generally accepted to be a monophyletic group, and within the Myrmeleontoidea, the antlions' closest living relatives are thought to be the owlflies (Ascalaphidae). A 2019 study finds Myrmeleontidae to be monophyletic, aside from Stilbopteryginae and Palparinae, which form separate clades closer to Ascalaphidae. The predatory actions of the larvae have attracted attention throughout history and antlions have been mentioned in literature since classical times. (Full article...)
Puzzled by the geographical distribution of wildlife and fossils he collected on the voyage, Darwin began detailed investigations and, in 1838, devised his theory of natural selection. Although he discussed his ideas with several naturalists, he needed time for extensive research, and his geological work had priority. He was writing up his theory in 1858 when Alfred Russel Wallace sent him an essay that described the same idea, prompting immediate joint submission of both their theories to the Linnean Society of London. Darwin's work established evolutionary descent with modification as the dominant scientific explanation of diversification in nature. In 1871, he examined human evolution and sexual selection in The Descent of Man, and Selection in Relation to Sex, followed by The Expression of the Emotions in Man and Animals (1872). His research on plants was published in a series of books, and in his final book, The Formation of Vegetable Mould, through the Actions of Worms (1881), he examined earthworms and their effect on soil. (Full article...)
A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. The wasps do not constitute a clade, a complete natural group with a single ancestor, as bees and ants are deeply nested within the wasps, having evolved from wasp ancestors. Wasps that are members of the clade Aculeata can sting their prey.
The most commonly known wasps, such as yellowjackets and hornets, are in the family Vespidae and are eusocial, living together in a nest with an egg-laying queen and non-reproducing workers. Eusociality is favoured by the unusual haplodiploid system of sex determination in Hymenoptera, as it makes sisters exceptionally closely related to each other. However, the majority of wasp species are solitary, with each adult female living and breeding independently. Females typically have an ovipositor for laying eggs in or near a food source for the larvae, though in the Aculeata the ovipositor is often modified instead into a sting used for defense or prey capture. Wasps play many ecological roles. Some are predators or pollinators, whether to feed themselves or to provision their nests. Many, notably the cuckoo wasps, are kleptoparasites, laying eggs in the nests of other wasps. Many of the solitary wasps are parasitoidal, meaning they lay eggs on or in other insects (any life stage from egg to adult) and often provision their own nests with such hosts. Unlike true parasites, the wasp larvae eventually kill their hosts. Solitary wasps parasitize almost every pest insect, making wasps valuable in horticulture for biological pest control of species such as whitefly in tomatoes and other crops.
Wasps first appeared in the fossil record in the Jurassic, and diversified into many surviving superfamilies by the Cretaceous. They are a successful and diverse group of insects with tens of thousands of described species; wasps have spread to all parts of the world except for the polar regions. The largest social wasp is the Asian giant hornet, at up to 5 centimetres (2.0 in) in length; among the largest solitary wasps is a group of species known as tarantula hawks, along with the giant scoliid of Indonesia (Megascolia procer). The smallest wasps are solitary parasitoid wasps in the family Mymaridae, including the world's smallest known insect, with a body length of only 0.139 mm (0.0055 in), and the smallest known flying insect, only 0.15 mm (0.0059 in) long. (Full article...)
It had several unique characteristics that differentiated it from many other arthropods, such as its long parabolic (nearly U-shaped) prosoma (head), its elongated first and second segments and the presence of paired "ridges" in the surface of its third to tenth tergites (dorsal halves of the segments). Furthermore, the opisthosoma (the "trunk") of Borchgrevinkium was triangular, and its telson (the posteriormost division of its body), short and wedge-shaped. It was a small animal, approximately 3 centimetres (1.2 inches) long.
The only known specimen of the genus was collected in 1956 and described in 1959 by the Russian paleontologistNestor Ivanovich Novojilov. He determined that it was a eurypterid and classified it in the familyMycteroptidae. However, its classification changed repeatedly over the years, being transferred from Eurypterida to Xiphosura and back to Eurypterida some time later. Borchgrevinkium is tentatively considered as a prosomapod, with more fossil material needed to ensure its current classification. (Full article...)
Chamberlin was a prolific taxonomist who named over 4,000 new animal species in over 400 scientific publications. He specialized in arachnids (spiders, scorpions, and relatives) and myriapods (centipedes, millipedes, and relatives), ranking among the most prolific arachnologists and myriapodologists in history. He described over 1,400 species of spiders, 1,000 species of millipedes, and the majority of North American centipedes, although the quantity of his output was not always matched with quality, leaving a mixed legacy to his successors. He also did pioneering ethnobiological studies with the Goshute and other indigenous people of the Great Basin, cataloging indigenous names and cultural uses of plants and animals. Chamberlin was celebrated by his colleagues at the University of Utah, however he was disliked among some arachnologists, including some of his former students. After retirement he continued to write, publishing on the history of education in his home state, especially that of the University of Utah.
Chamberlin was a member of the Church of Jesus Christ of Latter-day Saints (LDS Church). In the early twentieth century, Chamberlin was among a quartet of popular Mormon professors at Brigham Young University whose teaching of evolution and biblical criticism resulted in a 1911 controversy among University and Church officials, eventually resulting in the resignation of him and two other professors despite widespread support from the student body, an event described as Mormonism's "first brush with modernism". (Full article...)
Scale insects are small insects of the orderHemiptera, suborder Sternorrhyncha. Of dramatically variable appearance and extreme sexual dimorphism, they comprise the infraorder Coccomorpha which is considered a more convenient grouping than the superfamily Coccoidea due to taxonomic uncertainties. Adult females typically have soft bodies and no limbs, and are concealed underneath domed scales, extruding quantities of wax for protection. Some species are hermaphroditic, with a combined ovotestis instead of separate ovaries and testes. Males, in the species where they occur, have legs and sometimes wings, and resemble small flies. Scale insects are herbivores, piercing plant tissues with their mouthparts and remaining in one place, feeding on sap. The excess fluid they imbibe is secreted as honeydew on which sooty mold tends to grow. The insects often have a mutualistic relationship with ants, which feed on the honeydew and protect them from predators. There are about 8,000 described species.
The oldest fossils of the group date to the Early Cretaceous, preserved in amber. They were already substantially diversified by this time suggesting an earlier origin during the Triassic or Jurassic. Their closest relatives are the jumping plant lice, whiteflies, phylloxera bugs and aphids. The majority of female scale insects remain in one place as adults, with newly hatched nymphs, known as "crawlers", being the only mobile life stage, apart from the short-lived males. The reproductive strategies of many species include at least some amount of asexual reproduction by parthenogenesis.
Some scale insects are serious commercial pests, notably the cottony cushion scale (Icerya purchasi) on Citrus fruit trees; they are difficult to control as the scale and waxy covering protect them effectively from contact insecticides. Some species are used for biological control of pest plants such as the prickly pear, Opuntia. Others produce commercially valuable substances including carmine and kermes dyes, and shellac lacquer. The two red colour-names crimson and scarlet both derive from the names of Kermes products in other languages. (Full article...)
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1875 cartoon by Henry Worrall depicting Kansas farmers battling giant grasshoppers.
The Locust Plague of 1874, or the Grasshopper Plague of 1874, occurred when hordes of Rocky Mountain locusts invaded the Great Plains in the United States and Canada. The locust hordes covered about 2,000,000 square miles (5,200,000 km2) and caused millions of dollars' worth of damage. The swarms were so thick that they could cover the sun for up to six hours.
Efforts were made to stop the infestation, including eating the locusts. Following the plague, the population of Rocky Mountain locusts continued to decline each year after 1874 and in spring 1875, many of the hatched locust eggs died due to frost, contributing to their eventual extinction. (Full article...)
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Hemilepistus reaumuri is a species of woodlouse that lives in and around the deserts of North Africa and the Middle East, "the driest habitat conquered by any species of crustacean, not including insects which are now known to be crustaceans pancrustacea". It reaches a length of 22 mm (0.87 in) and a width of up to 12 mm (0.47 in), and has seven pairs of legs which hold its body unusually high off the ground. The species was described in the Description de l'Égypte after the French Campaign in Egypt and Syria of 1798–1801, but was first formally named by Henri Milne-Edwards in 1840 as Porcellio reaumuri. It reached its current scientific name in 1930 after the former subgenus Hemilepistus was raised to the rank of genus.
H. reaumuri occurs at great population densities and fills an important niche in the desert ecosystem. It feeds on plant leaves, obtains most of its water from moisture in the air and sand, and is in turn an important prey item for the scorpionScorpio maurus. H. reaumuri is only able to survive in such arid conditions because it has developed parental care of its offspring. Adults dig burrows which are inhabited by family groups, which are recognised using pheromones. The burrows are 40–50 cm (16–20 in) deep, and the woodlice retreat to the relatively cool and moist conditions of the burrow when surface conditions are unfavourable. The territorial limit of each colony is marked with a faecal embankment. (Full article...)
Aptostichus barackobamai (also known as the Barack Obama trapdoor spider) is a large species of trapdoor spider in the familyEuctenizidae named after the 44th President of the United States, Barack Obama. The species was first reported by Jason Bond of Auburn University in December 2012 as one of 33 new species of the genusAptostichus. The species is endemic throughout northern California, forming burrows out of silk, dirt, and sand, from which it attacks prey. This spider is identified by its dark red to red-brown coloration with a stripped or chevron patterned abdomen. Breeding occurs during the winter and females tend to be larger than their male counterparts. A. barackobamai is part of the larger Aptostichus icenogleispecies complex, and is a close relative to Aptostichus icenoglei and Aptostichus isabella. (Full article...)
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Alice E. Gray (June 7, 1914 – April 27, 1994) was an American entomologist and origamist. She worked as an entomologist at the American Museum of Natural History (AMNH) in New York for 43 years, writing, illustrating, and creating large models of insects. Known as the "Bug Lady", she conducted outreach and education in the museum, in local schools, and appeared on The Tonight Show. She began practicing origami first as an extension of her interest in insects, starting a tradition of using origami creatures to decorate the museum's Christmas tree. In the 1960s, she became more involved with the origami community and, in 1978, co-founded the Friends of the Origami Center of America in New York with Lillian Oppenheimer and Michael Shall, now known as OrigamiUSA. (Full article...)
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Two wingless adults and a nymph
The black bean aphid (Aphis fabae) is a small black insect in the genus Aphis, with a broad, soft body, a member of the orderHemiptera. Other common names include blackfly, bean aphid, and beet leaf aphid. In the warmer months of the year, it is found in large numbers on the undersides of leaves and on the growing tips of host plants, including various agricultural crops and many wild and ornamental plants. Both winged and wingless forms exist, and at this time of year, they are all females. They suck sap from stems and leaves and cause distortion of the shoots, stunted plants, reduced yield, and spoiled crops. This aphid also acts as a vector for viruses that cause plant disease, and the honeydew it secretes may encourage the growth of sooty mould. It breeds profusely by live birth, but its numbers are kept in check, especially in the later part of the summer, by various predatory and parasitic insects. Ants feed on the honeydew it produces, and take active steps to remove predators. It is a widely distributed pest of agricultural crops and can be controlled by chemical or biological means. In the autumn, winged forms move to different host plants, where both males and females are produced. These mate and the females lay eggs which overwinter. (Full article...)
Afraflacilla braunsi is a species of jumping spider in the genusAfraflacilla. First found in South Africa, the spider was subsequently observed living in Saudi Arabia, Turkmenistan, the United Arab Emirates and Yemen, although it is likely to have a wider distribution. First described in 1903 by George and Elizabeth Peckham, it was originally allocated to the genus Pseudicius with the name Pseudicius braunsii. After being renamed Icius braunsi in 1987, it was finally given its current name by Jerzy Prószyński in 2017. Pseudicius tripunctatus, now called Afraflacilla tripunctatus, is a synonym.
A small to medium-sized spider, Afraflacilla braunsi has a carapace that is between 1.7 and 1.9 mm (0.07 and 0.07 in) long and an abdomen 2.2 and 3.2 mm (0.09 and 0.13 in) long. The carapace is brown, sometimes with yellow sides. The abdomen is light brown or fawn with lighter yellow patches. It has large brown front legs, the remainder generally yellow and less robust. It makes noises by rubbing its forelegs against small hairs under its eyes. The copulatory organs are distinctive for the species. The male has a characteristically bulbous palpal bulb and a single long projection, or apophysis, extending from the palpal tibia. The female has very long and coiled insemination ducts that lead to large spermathecae. (Full article...)
Pseudastacus (meaning "false Astacus", in comparison to the extant crayfish genus) is an extinct genus of decapod crustaceans that lived during the Jurassic period in Europe, and possibly the Cretaceous period in Lebanon. Many species have been assigned to it, though the placement of some species remains uncertain and others have been reassigned to different genera. Fossils attributable to this genus were first described by Georg zu Münster in 1839 under the name Bolina pustulosa, but the generic name was changed in 1861 after Albert Oppel noted that it was preoccupied. The genus has been placed into different families by numerous authors, historically being assigned to Nephropidae or Protastacidae. Currently, it is believed to be a member of Stenochiridae.
Reaching up to 6 cm (2.4 in) in total length, Pseudastacus was a small animal. Members of this genus have a crayfish-like build, possessing long antennae, a triangular rostrum and a frontmost pair of appendages enlarged into long and narrow pincers. Deep grooves are present on the carapace, which is around the same length as the abdomen. The surface of the carapace is usually uneven, with either small tubercles or pits. Sexual dimorphism is known in P. pustulosus, with the pincers of females being more elongated than those of the males. There is evidence of possible gregarious behavior in P. lemovices in the form of multiple individuals preserved alongside each other, possibly killed in a mass mortality event. With the oldest known record dating to the Sinemurian age of the Early Jurassic, and possible species surviving into the Cenomanian stage of the Late Cretaceous, Pseudastacus has a long temporal range and was a widespread taxon. Fossils of this animal were first found in the Solnhofen Limestone of Germany, but have also been recorded from France, England and Lebanon. All species in this genus lived in marine environments. (Full article...)
... that adult males of the parasiticwaspEncarsia perplexa can only develop when a virgin female lays eggs in a fully developed larva of her own species?
Image 3Mature queen of a termite colony, showing how the unsclerotised cuticle stretches between the dark sclerites that failed to stretch as the abdomen grew to accommodate her ovaries (from Arthropod exoskeleton)
Image 4Ghost crab, showing a variety of integument types in its exoskeleton, with transparent biomineralization over the eyes, strong biomineralization over the pincers, and tough chitin fabric in the joints and the bristles on the legs (from Arthropod exoskeleton)
Image 14Formation of anterior segments across arthropod taxa based on gene expression and neuroanatomical observations, Note the chelicera(Ch) and chelifore(Chf) arose from somite 1 and thus correspond to the first antenna(An/An1) of other arthropods. (from Chelicerata)
Image 18In honeypot antrepletes, the abdomens of the workers that hold the sugar solution grow vastly, but only the unsclerotised cuticle can stretch, leaving the unstretched sclerites as dark islands on the clear abdomen (from Arthropod exoskeleton)
Image 26Some of the various hypotheses of myriapod phylogeny. Morphological studies (trees a and b) support a sister grouping of Diplopoda and Pauropoda, while studies of DNA or amino acid similarities suggest a variety of different relationships, including the relationship of Pauropoda and Symphyla in tree c. (from Myriapoda)
Image 27Decapods, from Ernst Haeckel's 1904 work Kunstformen der Natur (from Crustacean)
Image 31 Honeybee larvae have flexible but delicate unsclerotised cuticles. (from Arthropod exoskeleton)
Image 32This Zoea-stage larva is hardly recognisable as a crab, but each time it sheds its cuticle it remodels itself, eventually taking on its final crab form (from Arthropod exoskeleton)
Image 33 This fully-grown robber crab has tough fabric forming its joints, delicate biomineralized cuticle over its sensory antennae, optic-quality over its eyes, and strong, calcite-reinforced chitin armouring its body and legs; its pincers can break into coconuts (from Arthropod exoskeleton)
Image 34The house centipedeScutigera coleoptrata has rigid sclerites on each body segment. Supple chitin holds the sclerites together and connects the segments flexibly. Similar chitin connects the joints in the legs. Sclerotised tubular leg segments house the leg muscles, their nerves and attachments, leaving room for the passage of blood to and from the hemocoel (from Arthropod exoskeleton)
The cryptic mantis (Sibylla pretiosa) is a mantid native to southern Africa. Its common name comes from its ability to grow asymmetrically to match the vegetation of its environment. They have unusual leaf-like projections on the joint of their four walking legs and leaf-like wings, generally the only green portion of the insect's otherwise brown and mottled exoskeleton. Adult females grow to 5–6 cm (2.0–2.4 in) in length while the males are generally about 1 cm smaller.
Xylotrupes socrates (Siamese rhinoceros beetle, or "fighting beetle"), male, on a banana leaf. This scarab beetle is particularly known for its role in insect fighting in Northern Laos and Thailand.
The citrus root weevil (Diaprepes abbreviatus) is a major agricultural pest weevil in several Caribbean countries and the U.S. state of Florida. The larvae feed on the roots of the host plant for several months. They often eat the taproot of the plant, which can kill it by depriving it of water and nutrients or by making it vulnerable to infection by fungi, or water moulds.
Papilio polymnestor, the blue Mormon, is a species of swallowtail butterfly found in southern India and Sri Lanka. It is a woodland species, often seen on forest paths and near streams. The larvae feed on trees in the family Rutaceae, such as citrus. Young larvae are green with white markings and position themselves on the upper surface of leaves, relying on their cryptic colouring, which resembles bird droppings, for protection. Older larvae seek less conspicuous locations, and have a unique habit of securing their balance by weaving silk on the substratum. This adult male P. polymnestor butterfly was photographed in the Indian state of Kerala.
A Eusthenia species of stonefly. The order contains almost 3,500 known species, including the only known insects that are exclusively aquatic from birth to death. Stoneflies are believed to be one of the most primitive groups of Neoptera and are found worldwide, with the exception of Antarctica.
A compound eye is a visual organ found in certain arthropods. The compound eye consists of between 12 and 1,000 ommatidia, little dark/bright sensors. The image perceived by the arthropod is "recalculated" from the numerous ommatidia which point in slightly different directions. In contrast to other eye types, there is no central lens or retina. Though the resulting image is poor in resolution, it can detect quick movements and, in some cases, the polarization of light.
Dragonflies have about 30,000 facets to their compound eyes, giving them nearly a 360° field of vision.
An assassin bug belonging to the Reduviidae family of insects. A predatory insect so named because of its tendency to wait in ambush for its prey, the assassin bug uses its long rostrum to inject a lethal saliva that liquefies the internal structures of the prey, which are then sucked out.
A male Roesel's bush-cricket (Metrioptera roeseli), a European bush-cricket named after August Johann Rösel von Rosenhof, a German entomologist. Its song is very similar to that of Savi's Warbler. Its body length as an adult insect is 15 to 18 mm. It is brown with a pale margin to the sides of the pronotum. Its forewings usually reach midway along its abdomen at rest. However there is a macropterous form of this insect (f. diluta), in which the wings reach beyond the tip of the abdomen. This form appears predominantly during hot summers and enables the species to extend its geographical range rapidly while conditions are suitable; such migrations may also be in response to local overpopulation.
The orb-weaver spiders (family Araneidae) are the familiar builders of spiral wheel-shaped webs often found in gardens, fields and forests. The family is a large one, including over 2800 species in over 160 genera worldwide, making it the third largest known (behind Salticidae and Linyphiidae). The web has always been thought of as an engineering marvel.
Two Eastern Lubber grasshoppers (Romalea guttata) mating. Native to the southeastern and south central portion of the United States, it is well known both for its size and its unique coloration. During reproduction, the male grasshopper introduces sperm into the ovipositor through its aedeagus (reproductive organ), and inserts its spermatophore, a package containing the sperm, into the female's ovipositor.
The light blue soldier crab (Mictyris longicarpus) inhabits beaches in the Indo-Pacific region. Soldier crabs filter sand or mud for microorganisms. They congregate during the low tide, and bury themselves in a corkscrew pattern during high tide, or whenever they are threatened.
The Ozyptila praticola species of crab spider is found throughout Europe and the Middle East. They do not build webs to trap prey, but are active hunters. Crab spiders are so named because of their first two pairs of legs, which are held out to the side giving them a crab-like appearance. Also, like crabs, these spiders move sideways and backwards more easily than forwards.
Leptosia nina, known as the psyche, is a species of butterfly in the family Pieridae (the sulphurs, yellows and whites), found in the Indian subcontinent, southeastern Asia, and Australia. It has a small wingspan of 2.5 to 5 cm (1 to 2 in). The upper side of the otherwise white forewing has a large, somewhat pear-shaped, black spot; this spot is also present on the underside which is scattered with greenish dots and speckles, sometimes arranged in bands. This L. nina butterfly was photographed in Kerala, India.
The monarch butterfly (Danaus plexippus) is one of the best-known species of butterfly. Its wings feature an easily recognizable orange and black pattern, with a wingspan of 8.9–10.2 centimetres (3½–4 in).
Attacus taprobanis is a species of moth in the family Saturniidae native to southern India and Sri Lanka. This adult male, photographed in Kadavoor, Kerala, developed from a larva feeding on a mahogany tree. When ready to pupate, the larva formed a papery cocoon 7.5 cm (3 in) long interwoven with a leaf; before doing this, the larva had attached the leaf to the stem with a silken thread and cut the leaf stalk. The colours of the dying leaf provided camouflage for the pupa, and the adult insect emerged some 24 days later.
Compound eyes on a blue bottle fly. Unlike simple eyes, which have a single concave photoreceptive surface, compound eyes consist of a number of individual lenses (called ommatidia) laid out on a convex surface; this means that they point in slightly different directions. Compound eyes provide a wide field of view and can detect fast movement, but have low resolution.
A macro shot of the head of a dragonfly, focusing on its compound eyes. Dragonfly eyes have up to 30,000 facets; each one is a separate light-sensing organ or ommatidium, arranged to give nearly a 360° field of vision.
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